October 13, 2016
Plant Talk 10: Woody Plants
Hello plant enthusiasts…
Recent Events
Since the last class i have kept very very busy. The biggest adventure included teaching and a botanical survey at the SOURCE farm in St. Thomas Jamaica. It felt like a beneficial situation for everybody involved. i took note of over 100spp. On the land within the week i was there along with developing an ethnobotanical plant checklist of over 300spp. i was amazed to find out that of the roughly 3,000 spp. in Jamaica close to 900 are endemic and grow nowhere else in the world!
Returned home and the following day went right into a conference at Western Carolina University called Rooted in the Mountains focused on the Cherokee. Words can not express the sheer awesomeness and caliber of the presentations there. Unfortunately, attendance was low and it may or may not happen in this iteration again. However, i look forward to working more with colleague Dr. Jim Veteto and other speakers present like Tom Belt as well as Dr. Dave Cozzo and other folks to continue to have events in a similar vein in the future.
Right from that conference i went off to a workshop at Wellspring Mountain with herbalist Robert Rogers one of the North American authorities on Medicinal Mushrooms (R. D. Rogers, 2014b, 2014a; R. Rogers & Wasser, 2011). Then i taught botany following the workshop in exchange for my attendance.
Next onto a class for Corey Pine Shane’s Blue Ridge School of Herbal Medicine on Beverage Fermentation followed by some work at Warren Wilson College and checking in with my adopted grandmother Elly and then onto teaching at the legendary Jim Duke’s Green Farmacy north of Washington DC.
From there i went on to work in Frank Cook’s archives at the home of his father in Bethesda, MD. We are preparing for leading a trip to Peru through Plants and Healers International. As part of the preparation for the trip we are editing Frank’s book the Plants and Healers of Peru and Equador. His copious journals may help inform this process.
Since then i have been Guiding with No Taste Like Home, facilitated a Forage and Feast at Souflower Botanical Sanctuary and an Everything Apples event at the Appalachian Institute for Mountain Studies.
By far though the elephant in a rather crowded work room has been my time spent on behalf of the Asheville Tree Comission, Urban Agriculture Alliance and Buncombe county Master Gardener Cooperative Extension program largely focused on the mistake that led to the cutting down of nearly 20 year old apple trees in one of the first modern age edible parks in the country! The George Washington Carver edible park is supposed to be managed in partnership with the non-profit Bountiful Cities and the Buncombe County Fruit and Nut club with the oversight of entities like the Tree Commission. This unfortunate tree cutting event happening is especially disheartening as checks and balances were supposed to be built into the system! The action also goes in direct conflict with the work supported by the city at the Asheville Buncombe Food Policy Council and other agreements the city has made regarding planting edibles and promoting their growth in public places. There was a lot of citizen uproar and the action at the park was featured on the local TV news WLOS. This park has gotten press from many directions over the years from internationally known publications like the New York Times and Huffington Post to our own local Asheville Citizen Times and Mountain Xpress. Some historic photos and overview can be accessed at the Community Food Forests website. This park is one of the things that inspired me to move to Asheville in the late 90s as a young permaculturalist who dreamed of community food forests. It was great to see such a space already established right downtown. i sincerely hope that the various stakeholder groups in our local food movement can get on the same page to actualize the aspirations so many now hold dear.
Upcoming Events
In the next few weeks i will be teaching at the NC Arboretum, LEAF festival, and a Forage and Feast and Fermentation Extravaganza at Pickard’s Mountain in Chapel Hill, NC. Go to the Botany Every Day events tab on the website to learn more details. Hope to see some of you all around!
What’s Blooming
Most of what is blooming is in the Asteraceae including Boneset (Eupatorium perfoliatum), Goldenrod (Solidago spp.), Joe Pye Weed (Eutrochium purpureum), Ironweed (Vernonia), Marigolds (Tagetes spp.) Quickweed (Galinsoga) white Wood Aster (Eurybia) and Wingstem (Verbesina spp.). Sigrid Neilsen has authored a great guide to keying out the Asteraceae with lavish illustrations updated taxonomy and a free key and affordable price for the book which can all be found here.
Many annuals from the Lamiaceae are blooming as well including Basil (Ocimum spp.), Oregano, Pineapple sage (Salvia elegans), Russian Sage (Perovskia atriplicifolia) and others.
Other family representatives blooming include Knotweeds (Polygonum spp.) (Polygonaceae), Butterfly bush (Buddleja sp.) Scrophulariaceae, Globe Amaranth (Gompherena) (Amaranthaceae) and Hollyhocks (Alcea sp.) Malvaceae.
That said here in southern Appalachia the first killing frost is right around the corner. Leaves are changing colors and i have noticed Kudzu nipped by cold already. Time to bring in the house plants and plant some garlic!
Harvest
Such a time of abundance right now ready to be harvested! Typical things such as Corn, Beans, Squash, Tomatoes, Peppers, Potatoes and the like are in full effect from the garden. Choice wild edible greens like Lambsquarters (Chenopodium album) and Pigweed (Amaranthus spp.) are ready too. The big story though is the cultivated and wild fruit! Apples, Blueberries, Pears, Persimmons and Sumacs (Rhus spp.) are all booming around me. Soon Hawthorne (Crataegus spp.), American Rowan (Sorbus americana), and edible Viburnum spp. will be ready at higher elevations above 4,000 ft elevation. What is ready to harvest for thee?
Major Tree/Woody Plant Families Occurring around Appalachia and other Temperate Places
The focus of the current class is woody plants. The study of trees is called dendrology (Hardin, Leopold, & White, 2001). Silviculture is a rather academic term and practice that deals with the raising of trees often times for lumber purposes (Jose, Jokela, & Miller, 2007; Nyland, 2007; Puettmann, 2008; Smith, Larson, Kelty, & Ashton, 1996; G. Williams, 1979). I have recently taken to studying more intensively the neotropics of which i have come across two reference for Trinidad and Tobago and one for Costa Rica in particular regarding Silviculture (Jiménez Madrigal, 2002; R. C. Marshall, 1939; Reginald Charles Marshall, Trinidad, & Forest Department, 1930). A number of publications are available geared more towards landowners while often taking wider management concerns into account (Beattie & Levine, 1993; Hansen, Severson, & Waterman, 2011; Hilts & Mitchell, 2009; Shirley & Graves, 1967). The state of North Carolina offers great prices through their forestry program for native trees available to purchase by landowners. What kind of resources does your state or country offer?
Many people are aware of the tragedy of industrial forestry often employing the technique of clearcutting (Devall, 1993; Jensen & Draffan, 2003; Lansky, 1992). Here are a few resources that speak to the human aspects of deforestation in the tropics (“Palm Oil and Tropical Deforestation,” n.d.; Sponsel, Headland, & Bailey, 1996; Zarin, Alavalapati, Putz, & Schmink, 2004). Palm Oil deforestation in Indonesia and Central America is a particularly worrying. Alternatives for sustainable management of tropical forests are an important subject for discussion (Ford & Nigh, 2015; Southgate, 1998; Zarin et al., 2004). Of course there are many other resources regarding these issues. Which ones are you aware of or can you find?
Much of the information below comes from my graduate research on the plants of Appalachia like many of the classes this year. However, it has since been expanded to cover information about woody plants from other temperate areas and a bit about subtropical/tropical areas as well. Here are featured 22 of the approximately one hundred and sixty plant families occurring in the Blue Ridge part of Appalachia (Wofford, 1989). The focus is on families that include a majority of woody species. Families are listed here in monographs because they have members representing a multitude of uses across a broad spectrum of topics. All families are listed alphabetically by botanical name. A common name corresponding to the typic genus for the family accompanies the title. In the case of the Aquifoliaceae and Caprifoliaceae there is no corresponding typic genus known and another common name has been chosen.
Due to time constraints some notable families that are useful or make up a sizable part of the Appalachian woody flora have been excluded. These include Witchhazel (Hamamelidaceae), Mulberry (Moraceae), Olive (Oleaceae), and Elm (Ulmaceae). The references consulted for the statistics of family geographic representation include Flowering Plant Families of the World (Heywood, Brummitt, Culham, & Seberg, 2007), Guide to the Vascular Plants of the Blue Ridge (Wofford, 1989) and a Synonynized Checklist of the Vascular Flora of the United States, Canada and Greenland (Kartesz, 1994). The work by the Karteszs also covers Puerto Rico, Hawaii and the Virgin islands. Heywood et al. split some families up that are considered as larger in the older sources of Wofford and Kartesz. Notations are included in the discussion of taxonomy for individual families. Several other books might be worth referencing for those interested in plant family patterns. (Castner, 2004; Evert, Eichhorn, & Raven, 2012; W. S. Judd, Campbell, Kellog, Stevens, & Donahue, 2008; Zomlefer, 1994). Newer references i have come across more recently include (Group, 2009; Mabberley, 2008a; Simpson, 2010; Soltis, 2005; Spichiger, Savolainen, Figeat, & Jeanmonod, 2004; Weakley, 2015; Woodland, 2009). The Plant List is an excellent source of online taxonomy especially at the species level.
The role of trees is something important to consider. Pragmatically trees provide habitat, food, building materials, textiles, shade, paper, carbon sequestration and many more services as well. However, on an energetic note, certain trees have given me more solace than any person ever could. Trees have played many other spiritual and intangible roles in society as well (Altman, 2000; Caldecott, 1993; Hageneder, 2005; Hugo, 2011; Wells, 2010). Old growth forests in particular tend to very inspiring (Bash, 2002; Kelly & Braasch, 1988; Maloof, 2011; Rapp, 2002; Valentine, 2011). The history of trees in general is rather fascinating (Thomas, 2014; Wells, 2010).
We have a couple great resources for the role of woody plants in supporting wildlife in the southeast (Huegel, 1995; J. H. Miller & Miller, 2005). There is also another good framing of this subject from a permaculture perspective(Jacke & Toensmeier, 2005a, 2005b). The difference between native species and exotic ones in this regard can be profound (Tallamy, 2009) Oaks and Cherries are some real winners in particular in regards to supporting wildlife. Many microbial associations are supported by various trees as well. Forestry Professor Suzanne Simard at the University of British Columbia has done a lot to highlight the ways trees communicate to each other in part mediated by microbial connections.
Many people think of Native Americans as passive in their influence on the forest. However, a literature has developed that shows a much more active role in managing arboreal ecosystems (Abrams & Nowacki, 2008; Black, Ruffner, & Abrams, 2006; Delcourt & Delcourt, 1998; Delcourt, Delcourt, Ison, Sharp, & Gremillion, 1998; McEwan, Dyer, & Pederson, 2011).
Trees in urban environments have very particular needs and constraints (E. F. Gilman, 1997; Watson, 1998). Gingko biloba is a popular addition to cityscapes as is the London Plane tree (Platanus x acerifolia). Elms were a historical favorite that was unfortunately afflicted by the Dutch Elm disease (Ophiostoma ulmi) a few decades ago and lost to the cityscape for that reason. Diane Rehm hosted a great show on Urban Trees. Tree planting and care is an art that may take much study to master (E. Gilman, 2001; Watson, 1994, 2005). Selecting the right tree for the landscape can be aided by a number of good resources as well (Dirr, 1997, 1998; Flint, 1997; Tennessee Valley Authority, 1998). One aspect that people sometimes select for is variegation (Houtman, 2004). DAVEY is one operation with over 130 years of experience in arboriculture. There is a robust literature documenting the benefits and ecosystems services of urban forests (Dwyer, Nowak, Noble, & Sisinni, 2000; Escobedo & Nowak, 2009; Gregory McPherson, 1992; Jonnes, 2016; McHale, Gregory McPherson, & Burke, 2007; E. G. McPherson et al., n.d.; E. G. McPherson & Simpson, 1999, 2003; G. E. McPherson, Nowak, & Rowntree, 1994; G. McPherson, Simpson, Peper, Maco, & Xiao, 2005; Nowak et al., 2008; Nowak & Crane, 2000; Nowak, Crane, & Stevens, 2006) .
Edible landscaping is becoming more and more popular these days (Creasy, 1982; M. Judd, 2013; Kourik & Kane, 1986; Tiruneh & Hager, 2011). One type of edible landscaping dealing with trees in particular is called Forest Gardening (Crawford, 2010; Hart, 1996; Jacke & Toensmeier, 2005a, 2005b; Mudge, Gabriel, & Munsell, 2014; Whitefield, 1998). Village Homes outside of Davis, CA is one excellent example of forest gardening being implemented on community scale (Corbett & Corbett, 2000). A couple resources cover many of the nuts of the world and in the eastern US cultivated and wild including woody plants and some non-woody ones as well (Krochmal & Krochmal, 1982; Menninger, 1977).
Conifers vs Broadleaves
One of the biggest distinctions in the tree world is between hardwoods also known as broadleaves and conifers. Conifers are a more ancient lineage with a once wider distribution. A number of great references cover the amazing world of conifers (Eckenwalder, 2009; Farjon, 2008, 2010; Elbert L. Little, n.d.).
Conifers
The two main families of conifers include the Pinaceae and Cupressaceae. Pinaceae members tend to have needles in different numbers and arrays in place of broad leaves. Pine (Pinus), Fir (Abies), Spruce (Picea), Douglas Fir (Pseudotsuga), Hemlock (Tsuga), Classic Cedar (Cedrus) and Larch (Larix) are all included in this family. These genera also tend to have vitamin C in their needles which may come in handy in the case of being cut off from traditional sources. However, too much intake of Pinaceae needle tea can be damaging due to the presence of resins (Elpel, 2013).
Cupressaceae members often have foliage with imbricate (layered) scales that are sharp to the touch. The most common members are Juniper (Juniperus) and Cypress (Cupressus). Many trees that formerly went by Cupressus in the Americas have changed to the genus (Callitropsis) (Judd et al., 2008). Thuja and Chamaecyparis occur in the east and the west and go by a host of common names including types of Cedar. Incense Cedar (Calocedrus) is a common genus in the western United States. i am aware of a few book resources for the study of western trees specifically (Earle, Spellenberg, & Nelson, 2014; Elbert L. Little, 1976; Petrides, 1998b, 2005; Sudworth, 1967).
The formerly separate members of Taxodiaceae are now included within the Cupressaceae family (Judd et al., 2008). North American members include California Redwood (Sequoia sempervirens) Giant Sequoia (Sequoiadendron giganteum) and Bald Cypress (Taxodium spp.). Dawn Redwood (Metasequoia glyptostroboides) was a species thought to be extinct until it was found in the 1940s’ (Judd et al., 2008). This sub-family holds the tallest and biggest tree titles for the planet and also some of the oldest trees as well. Bald Cypress and Dawn Redwood also belie the title evergreen as they both lose their leaves in the winter. Larch from the Pinaceae is another example in this regard. Other more exotic genera traditionally included in the Taxodiaceae include Cryptomeria, Athrotaxis, Cunninghamia, Glyptostrobus, Sciadopitys, and Taiwania.
Smaller families of conifers include the Yew (Taxaceae), Podocarp (Podocarpaceae), Monkey Puzzle (Araucariaceae), Umbrella Pine (Sciadopitayaceae) and interestingly the not needled and monogeneric Ginkgo (Ginkgoaceae). The Podocarp and Monkey Puzzle families often occur in the Southern Hemisphere. One of the largest trees i have ever encountered was a Podocarpus centuries/millenia old growing near 10,000 ft elevation and the Cerro de la Muerte in Costa Rica. It is located on the Seeyle family land which i highly recommend visiting if you are ever in the area www.lasvueltas.com. It took over 15 people to make a ring around this massive elder tree and is one of my greatest memories from travels with Frank Cook.
In the northern temperate regions conifers tend to grow on more harsh sites. Some examples include dry, fire prone, windswept, acidic, snow buried or frigidly cold. They tend to have shallower roots than broadleaves, thicker bark and sometimes quicker growth as some adaptations for their chosen path. Most of them will not sprout from stumps. Redwoods are a notable exception. More conifers are put into plantations than probably any other trees. Pines in particular are popular in this regard i.e. Loblolly (Pinus taeda), Slash (Pinus elliotii) and Monterey (Pinus radiata).
The Hemlock Wooly adelgid (Adelges tsugae) and Balsam Wooly adelgids (Adelges piceae) are devastating pests. The Hemlock “Reishi” mushroom (Ganoderma tsugae) is one famous associated fungal species. It is a bit of a silver lining on the fact that most Hemlocks in the east have been decimated by the adelgid pest. An entomologist friend Richard McDonald recently told me that he has high hopes for beneficial insect identified in the western U.S, that may help control the hemlock wooly adelgid in the east as well. Pine bark beetles (Dendroctonus spp.) are another challenging insect pest. Pine blister rust (Cronartium ribicola), which is hosted on Currants (Ribes spp.) acts as a microbiological threat. Mushrooms from the family Boletaceae famously co-habitate with conifers in beneficial mycorrhizal relationships.
A lot of wild Ericaceae members are associated with the understory of conifers out west and to a certain degree in the east as well. Members of the non-photosynthesizing group formerly known as the Monotropaceae have now been subsumed by the Ericaceae and tend to grow along with conifers as well. The Rhododendrons within the Ericaceae have an extensive literature all their own (Bryant, 1996, 2001; K. Cox, 2005; K. N. E. Cox & Glen, 1998; P. A. Cox & Cox, 1988; Cullen, 2005; Davidian, 1982, 2003a, 2003b, 2003c; Hawthorne, 2014; Leach, 1961; Reilly, 1992; Salley & Greer, 2005; Towe, 2004; Vanveen, 1986; Young, 1981).
Hardwoods
Hardwoods tend to grow on nice sites and have tap roots. They are mostly deciduous in the north but often evergreen in the tropics. They often prefer more basic soils than conifers. Their diverse arrays of fruits also tie them into a wider web of fauna. Many specific broadleaf trees have whole industries built around them including examples like tool handles, pencils, toothpicks, baseball bats and hockey sticks.
Many of the broadleaf trees that are native to North America are currently under assault from exotic insect or disease pests (C. E. Little, 1995). Emerald Ash Borer (Agrilus planipennis) has killed millions of Ash trees and continues to spread. Gypsy Moths (Lymantria spp.) and Sudden Oak Death (Phytopthora ramorum) are affecting Oaks. Dogwood Anthracnose is taking out these iconic small trees. Other species are under attack as well including Butternut (Juglans cinerea) and Red Bay (Persea borbonia). Chestnut Blight (Cryphonectria parasitica) and Dutch Elm disease are afflictions of the last century that have radically changed forest composition in the east. The plight of tree diseases has been well documented (Brosi, 2010; Clark, Brosi, Schlarbaum, & Grissino-Mayer, 2008; Coyier & Roane, 1986; Dalgleish & Swihart, 2012; Dreistadt, 2004; Hepting, 1971; C. E. Little, 1995; Sinclair & Lyon, 2005).
Woody Plant Identification
Flowers or fruits are always the easiest way to identify a plant in combination with the rest of the characteristics. The first characteristic that i pay attention to elsewise is leaf orientation i.e. whether a plant is opposite or alternate. Most temperate world opposite leaved woody plants fall into the pneumonic device that i helped develop in collaboration with colleague and friend Robin Allison PrincessBBAAADCapSapSweetLove Pronounced Princess buh buh aaaaaaaad CapSapSweetLove
Princess Tree (Pawlowniaceae)
(B) ignoniaceae i.e. Cross Vine (Bignonia) Catalpa, Trumpet Creeper (Campsis)
(B) uxaceae i.e. Boxwood (Buxus) and Pachysandra
(A) sh i.e. Ash (Fraxinus), Jasmine (Jasminum) Olive (Olea), Privet (Ligustrum)
(A) doxaceae i.e. Elderberry (Sambucus) and Viburnum
(A) ucubaceae i.e. Japanese Laurel (Aucuba japonica)
(D) ogwood (Cornus)
(CAP) rifoliaceae i.e. Honeysuckle (Lonicera)
(Sap) indaceae i.e. Buckeye (Aesculus) Golden Rain Tree (Koelreuteria) and Maple (Acer).
(Sweet) shrub (Calycanthaceae)
(Love) Heart’s a bustin (Euonymous), Bittersweet (Celastrus), (Celastraceae)
Compound versus simple leaves are the next distinguishing trait with woody plant identification especially. Compound leaves are fairly uncommon and more distinguished depending on leaf orientation. Opposite compound typically denotes Mochatel (Adoxaceae), Ash (Oleaceae) and Maple/Buckeye (Sapindaceae) families. Woody plants with alternate compound leaves typically fall in the Walnut (Juglandaceae), Rose (Rosaceae) or Bean (Fabaceae) families. Tree of Heaven (Ailanthus altissima) (Simaroubaceae) is another pinnately compound alternate plant that is invasive in many areas especially in disturbed locales. However, like many invasive plants it is also a potent medicinal (Scott, 2010).
Leaf margin, hairiness, buds, and scars are additional traits used in dendrological identification. Hairiness denoted by such terms as hirsute, pillose, pubescent, stellate, strigose, tomentose, , etc…are often reason enough for classic species delineation. Winter identification mostly depends on bark, bud type (number/shape of scales) and leaf/vascular bundle scars. We have played winter tree id in the Asheville area to much enjoyment and delight as well as humbled bafflement even in familiar terrain. Some books exist that can help specifically with winter id (Core & Ammons, 1999; Gilkey & Packard, 2001; Harlow, 1946; Jones & Wofford, 2013; Lance, 2004; Symonds, 1958; Trelease, 1967; Watts & Watts, 1970; M. D. Williams, 2007; Wojtech, 2011). Trees and shrubs can provide quite a bit of “winter interest” in the garden when selected properly (Buffin, 2005; Price, 2007; Simeone, Curtis, Barnett, & Tankersley, 2005). My friend and teacher Sevensong has a great handout on winter tree id that he has graciously offered to share online.
Regional Guides
Many great references exist to help one in the exploration of woody plants and their uses (Brockman, 1986; Dirr, 1998; Elias, 1980; Hardin, Leopold, & White, 2001). In Appalachia the references i use most are represented by a handful of books (Petrides, 1998a; Radford, Ahles, & Bell, 1968; Swanson, 1994; Weakley, 2012; Wofford, 1989). Swanson in particular is key! Other older and less comprehensive references are also available (Stupka, 1964). A couple bibliographies have been developed for information on Southern hardwoods too (W. D. Miller, 1967, 1974).
In the Sierra Nevada’s of the U.S. i have been thankful for a compact lightweight book by Petrides (2005). In the past i have also consulted bigger more comprehensive resources for woody plants of the Western U.S. (Petrides, 1998b; Sudworth, 1967).. There is also a reference specific to trees and shrubs of California (Stuart & Sawyer, 2001). The Flora of the California is known as the Jepson Manual (Hickman, 1993). A newer version has been published as well (Baldwin, Goldman, Keil, Patterson, & Rosatti, 2012). California also has great online botany information through Cal Flora These are the definitive resources to the over 5,000 plant taxa that occur within the Golden state. Quite a few books have been written for exploration of woody plants in the desert Southwest (J. E. Bowers, 1998; J. Bowers & Wignall, 1993; Carter, 1997; Elmore, 1976; MacKay, 2003).
Specific resources have been generated for the more tropical clime of Florida (Kurz & Godfrey, 1962; Elbert L. Little, 1978; G. Nelson, 1998). i am aware of a few references for Costa Rica as well (Céspedes & Lindquist, 2007; Condit, Pérez, & Daguerre, 2010; Poveda, Jiménez, & Zamora, 2011; Zamora, Jimenez, & Poveda, 2004). There is a handy pocket guide for trees and shrubs of Hawaii as too (Pratt, 1998). Thanks to my recent travel and interest in the Caribbean i have come across quite a few resources on woody plants from the region (Leiva, 2007; Elbert Luther Little & Wadsworth, 1964; Marcelle, Hawthorn, & Jules, 2004; Pertchik, Pertchik, & White, 1951; Seddon & Lennox, 1980; Storer, 1964; Tanner, 1977).
i know of at least one good guide to trees of England (Sterry & Sterry, 2008). i also have a guide to both trees and shrubs from a company based in England that covers over 9,000 plants in over 650 genera (Nurseries, 1992)! One guide covers over 500 trees of both Europe and North America (Phillips, 1978). Another guide by Phillips (2002) references 450 genera of trees and shrubs once again accompanied by great photographs.
Some specific resources for shrubs often combined with vines are available as well (Curtis, 1945; Foote & Jones, 1989; Grimm, 1966; Hyams, 1965; Keeler, 1969; Tennessee Valley Authority, 1998; Zucker, 1966). My favorite resource by George Symonds (1963) contains photographs for side by side comparisons of different species bark, flowers and fruits.
A number of resources also cover trees of North America in a broad fashion (Brockman, 1986; Peattie, 2013; Russell & Cutler, 2003; Sargent, 1965). According to one such reference there are at least 652 native species and over 100 introduced species of trees in North America (Elias, 1980).
Overall guides to the trees of the world exists as well (Russell & Cutler, 2003; Russell & Walters, 2007). i also have come across one exquisite guide to trees of the world covering almost 2,000 species and cultivars with awesome illustrations and ethnobotanical references (More & White, 2005). One rather comprehensive partly digital resource covers 8,500 different trees and shrubs of the world and also come with a handy interactive CD (Wasson, 2001). A book by Thomas Pakenham (2003) takes note of remarkable trees of the world and makes for a jaw dropping bucket list for the intrepid traveler and pilgrim.
Clearly a lot of awesome resources are out there in book form and certainly some great websites exist as well. One great free resource that i recently came across with numerous publications on tree care is from Bartlett Tree Experts http://www.bartlett.com/resourceList.cfm.
Monographs
Below are monographs for most of the major woody plant families in the temperate world! Families are listed alphabetically by scientific name. Your support to develop this further is greatly encouraged. Factoids and funds could move this along to a more comprehensive place!
Family descriptions are covered systematically. The first section includes a chart showing distribution in the world, North America and the Blue Ridge using the sources underlined in the introduction above. A modern taxonomic commentary depending on three main sources follows (Heywood, Brummitt, Culham, & Seberg, 2007; W. S. Judd et al., 2008; Weakley, 2015). Next follows a listing of genera known in Appalachia according to Wofford (1989). The number of species for each genus is in parentheses. The first number is for native species. The second number when present is for introduced species. The introduced species numbers are also highlighted to further show their patterns of distribution. Introduced plants in general tend to be represented by one species from an exotic genus. The native species almost always out number introduced species when contained within the same genus. A list of taxa excluded by Wofford follows. Excluded taxa are normally reported from a limited range and are often cultivated plants that may escape locally. Within the above three botanical sections all plant names are in scientific terminology only. In the below usage categories common names precede scientific names at least the first time the plant is mentioned i.e. Eastern Hemlock (Tsuga canadensis). Common plant names are capitalized as usual because they are at least as important as people to me!
Finally, species that fit into the categories of food, medicine, poison, insect affiliation, invasiveness, dye plants, other crafts, aesthetics and aromatics are included. Foods included comprise wild and cultivated taxa. One source is the primary reference mixed with over a 15 years of personal experience regarding wild foods (Couplan, 1998). Don’t ever eat flowers or other parts of plants from a florist or other synthetic chemical intensive source. Always be absolutely positive of the identification and utilization of anything you plan to consume! Medicinal notations address only some extremely important plants due to time limitations. Invasiveness is mostly only covered relative to Appalachia and specifically inclusion in the Blue Ridge Flora. One cited reference was used primarily (J. H. Miller, Chambliss, & Bargeron, 2006). However, personal experience with exotic invasive plants informed commentary that is not cited. Many invasive plants in Appalachia are also invasive throughout their introduced range. Plant uses by bees were compiled primarily from three comprehensive book sources with the addition of 14 years of personal experience (H. B. Lovell, 1977; J. H. Lovell, 1999; Pellett, 1977). Dye plants included are noted when not so plentiful to be confusing. For the Fabaceae and Rosaceae for instance only some dye plants of special interest are included. Highlighted sections are still a work in progress.
Aceraceae/Maple Family (Now included in the Sapindaceae)
Blue Ridge |
|
North America |
|
World |
|
Genera |
1(n) |
Genera |
1 |
Genera |
|
Species and Lower taxa |
8(n) 1(i) |
Species |
19 |
Species |
|
|
|
Lower Taxa |
21 |
|
|
Taxonomic commentary: This family is now often included as a part of the more tropical Sapindaceae (Heywood et al., 2007; Judd et al., 2008; Weakley, 2008). That is why no distribution occurs for world scope.
Blue Ridge Genera: Acer
Food: Sap from various species. Seeds are also said to be edible (Couplan, 1998).
Insects: Provides early forage for bees in February/March
Invasives: Norway maple (Acer platanoides)
Dye Plants: Acer spp. (Eaton, 1973; Fern, 2009; A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis, 1967)
Other Crafts: Acer spp.
Aesthetics: Paper bark maple (Acer griseum) is a marvelous landscape tree with exfoliating bark. Many other “Japanese” maples i.e. A. palmatum etc. are popular landscape plants.
Anacardiaceae/Cashew Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2(n) |
Genera |
12 |
Genera |
82 |
Species |
7(n) |
Species |
37 |
Species |
700+ |
Lower taxa |
|
Lower Taxa |
22 |
|
|
Blue Ridge Genera: Toxicodendron (3), Rhus (4)
Food: Fruits of Sumac (Rhus spp.) have been used by Native Americans and others to make a type of pink lemonade due to malic acid. Cold infusion and straining is necessary. Heat brings out bitter tannins. Many popular exotic food stuffs are in this family. These include Pistachio (Pistacia spp.), Cashew (Anacardium occidentale), Mango (Mangifera indica), and Brazilian pepper (Schinus spp.). In the Caribbean the June Plum (Spondias sp.) is also enjoyed. A type of Sumac (Rhus coriaria) is ground up and used as a flavoring spice called Zaatar in the Middle East.
Toxicity: Poison Ivy (Toxicodendron radicans), Poison Oak (T. toxicarium), and Poison Sumac (T. vernix) can cause severe skin irritation for susceptible individuals. Poisonwood/Burn Plum (Metopium spp.) Allergic reactions are known to occur for certain people with many if not all the members of this family!
Insects: Bees freely work Rhus spp. for nectar.
Invasiveness: Poison ivy is native but can quickly take over disturbed areas.
Dye Plants: Smoketree (Cotinus coggygria), Sumacs (Rhus copallina, R. copallinum, R. coriaria, R. glabra, R. hirta, R typhina) (Bliss, 1993; Cannon & Cannon, 2003; Fern, 2008; Krochmal & Krochmal, 1974; Moerman, 1998; Nicholson & Clovis, 1967).
Aesthetics: Fruits of Rhus spp. make great additions to flower arrangements fresh or dry. Cotinus is a beautiful landscape plant with purple cultivars that can also be employed in flower arrangements to striking effect.
Aquifoliaceae/Holly Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2(n) |
Genera |
2 |
Genera |
1 |
Species |
4(n) |
Species |
29 |
Species |
400+ |
Lower taxa |
2 |
Lower Taxa |
9 |
|
|
Taxonomic commentary: Nemopanthus has now been moved to Ilex (Heywood et al., 2007).
Blue Ridge Genera: Ilex (4), Nemopanthus (1)
Food: The roasted leaves of some members have been used to prepare a caffeinated beverage including Yerba Mate (Ilex paraguariensis) and Yaupon Holly (I. vomitoria) (Couplan, 1998). Ilex cassine and I. guayusa also have significant amounts of caffeine (Edwards & Bennett, 2005; Wink & Van Wyk, 2008). The fruits of Gallberry (Ilex glabra) are said to be edible (Couplan, 1998).
Toxicity: Triterpine saponins and their esters (Wink & Van Wyk, 2008). The berries are often considered to not be edible, however, the leaves tend to be not toxic (Nelson, Shih, & Balick, 2007).
Insects: Gallberry (Ilex glabra) is important for bees in warmer areas (Pellett, 1976). Many other species of Ilex are good sources of nectar as well.
Wildlife: Many birds appreciate the fruits of Hollies.
Invasiveness: None known
Dye Plants: American Holly (Ilex opaca)
Other Crafts: Ilex spp. have traditionally been use for wood work.
Aesthetics: Many species are very attractive in the landscape for evergreen foliage and showy fruits. Plants are dioecious so males and females are required for fruiting.
Araliaceae/Spikenard Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2(n) 1(i) |
Genera |
11 |
Genera |
41-50 |
Species |
6(n) 1(i) |
Species |
49 |
Species |
1,450 |
Lower taxa |
|
Lower Taxa |
114 |
|
|
Blue Ridge Genera: Aralia (4), Hedera (1), Panax (2)
Food: The leaves and roots of Sarsaparilla (Aralia californica) are used by Daniel Nicholson in California for tea. This is the only native member of Araliaceae in the state. Other members of Aralia from the Eastern U.S. have been used similarly especially in the preparation of root beer (Peterson, 1978). Some young shoots of Aralia spp. have been eaten in the spring (Couplan, 1998). However, i know from direct experience that Aralia spinosa may cause some irritation especially if one tries to consume it raw.
Medicine: Multiple species of Panax are adaptogenic (Winston & Maimes, 2007; Yance, 2013) Aralia spp. may be medicinal as well. Devil’s Club (Oplopanax horridus) is a major medicinal in the northwest U.S. (Pojar et al., 1994).
Toxicity: Triterpine saponins occur in the Araliaceae including English Ivy (Hedera helix) (Wink & Van Wyk, 2008). Hedera helix may also cause contact dermatitis (Nelson et al., 2007). Members of Aralia may cause irritation to susceptible individuals especially if consumed raw.
Insects: Insects love members of this family in bloom.
Invasiveness: English ivy (Hedera helix) is a prolific exotic invasive.
Dye Plants: Hedera helix (Cannon, 1994; Fern, 2009).
Other Crafts: Vines of ivy can be used for various decorations.
Aesthetics: Hedera helix is famous for covering buildings but can cause some damage to the outside surface. Many Aralia spp. are beautiful plants with showy blossoms. Devil’s Club (Oplopanax horridus) is stunning in the understory of the Pacific Northwest conifer forests.
Miscellaneous: English Ivy is one of the best plants studied for improving indoor air quality (Wolverton, 1997).
Berberidaceae/Barberry Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
5 |
Genera |
10 |
Genera |
12-16 |
Species |
6(n) 1(i) |
Species |
35 |
Species |
650-700 |
Lower taxa |
|
Lower Taxa |
6 |
|
|
Blue Ridge Genera: Berberis (2), Caulophyllum (2), Diphylleia (1), Jeffersonia (1), Podophyllum (1)
This is a mostly vegetative family with one prominent shrubby member (Berberis) in Appalachia and a host of medicinal uses.
Food: The edible berries are sour and often need to be sweetened in the Barberry (Berberis) and cultivated Oregon grape (Mahonia) genera. Fruits of Mayapple (Podophyllum peltatum) may be consumed when fully ripe!!!
Medicine: Blue Cohosh (Caulophyllum thalictroides) is a potent medicine for women’s issues. Berberine contained in many members is a powerful immune boosting anti-microbial medicine. Podophyllotoxin from Mayapple is used as an anti-cancer drug (Duke & Foster, 1999).
Toxicity: This family tests the boundaries between food, medicine and poison. Blue Cohosh (Caulophyllum thalictroides) contains an alkaloidal convulsant poison (Nelson et al., 2007). Podophyllum contains a mitotic inhibitor (Nelson et al., 2007).
Insects: Berberis spp. are attractive to bees but rarely provide much honey.
Invasiveness: The exotic Japanese Barberry (Berberis thunbergii), Beale’s Barberry (Mahonia bealei) and Heavenly Bamboo (Nandina domestica) all can be invasive (Miller et al., 2006).
Dye Plants: Many species of Barberry (Berberis spp.) and Oregon Grapes (Mahonia spp.) can be used for dyes (Bliss, 1994; Cannon, 1994; Fern, 2009; A. Krochmal & C. Krochmal, 1974).
Aesthetics: The flowers of Oregon grape are often showy and slightly fragrant. The fruits of various Barberries and Oregon grapes are attractive. Though spiny, they can be used in flower arrangements.
Betulaceae/Birch Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
5 |
Genera |
5 |
Genera |
6 |
Species |
13(n) 1(i) |
Species |
32 |
Species |
130 |
Lower taxa |
2 |
Lower Taxa |
47 |
|
|
Blue Ridge Genera: Alnus (3), Betula (7), Carpinus (1), Corylus (2), Ostrya (1)
Food: Nuts from Corylus including the European Filbert (C. avellana) and the American Hazelnut (C. americana). The inner bark of the Alder has a tradition of use by Native Americans as a survival food (J. Pojar et al., 1994). Alder has also been used to smoke food. A number of Birch species can be tapped for their sap which can be consumed raw or brewed into various different types of beverages (Couplan, 1998). The tradition in Northern and Eastern Europe of using the saps for brewing has been studied in particular (Svanberg et al., 2012).
Medicine: Alder is a typical medicine for the Native people of the Pacific Northwest (J. Pojar et al., 1994). Black Birch (Betula lenta) contain methyl salicylates which act as mild analgesics akin to the chemistry in Aspirin and correspond to the smell of wintergreen (Duke & Foster, 2014).
Toxicity: ?
Insects: Alders (Alnus spp.) are an important early source of pollen for bees.
Invasiveness: None known
Dye Plants:
Alder (Alnus spp.) (Bliss, 1994; Eaton, 1973; A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis, 1967).
Birch (Betula spp.) (Cannon, 1994; Fern, 2009)
Other Crafts: Wood used by western tribes for bowls and masks (J. Pojar et al., 1994).
Aesthetics: Many species make handsome landscape trees especially Harry Lauder’s walking stick (Corylus avellana ‘contorta’). Purple leaved varieties have been developed as well.
Miscellaneous: Alders fix nitrogen though a different microbial association regarding actinomycetes rather than the better known Rhizobium and Bradyrhizobium from the Fabaceae. Wax Myrtle (Myrica gale) and Sweet Fern (Comptonia) from the Myricaceae and Ceonothus from the Rhamnaceae do this as well (Evert et al., 2012).
Caprifoliaceae/Honeysuckle Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
7 |
Genera |
11 |
Genera |
12 |
Species |
22(n) 4(i) |
Species |
82 |
Species |
250 |
Lower taxa |
2 |
Lower Taxa |
59 |
|
|
Taxonomic commentary: The Caprifoliaceae is a messy family in transition. Elderberries (Sambucus spp.) and Viburnums (Viburnum spp.) have been moved to Adoxaceae (Spears, 2006). A number of genera have sometimes been given their own families i.e. Sambucaceae, Diervillaceae, Linnaeaceae (Heywood et al., 2007). American Elderberries are sometimes treated as subspecies of the European one (Sambucus nigra) but our most recent southeastern flora keeps them at the level of species rank (Weakley, 2015).
Blue Ridge Genera: Diervilla (3), Linnaea (1), Lonicera (3n,3i), Sambucus (2), Symphoricarpos (2), Triosteum (3), Viburnum (9) Wofford excluded: Lonicera fragrantissima and Lonicera xylosteum
Food: Fruits from Black Elderberry (Sambucus canadensis) and Blue Elderberry (Sambucus cerulea) are used for jelly, wine, syrup etc. Some black fruited Viburnums are edible raw (V. cassinoides, V. alnifolia, V. lantana, V. nudum, V. prunifolium, V. rufidulum) and red fruited Viburnums are best cooked (V. trilobum, V opulus) (Couplan, 1998). Fruits from the horse gentian (Triosteum perfoliatum) can be roasted as a coffee substitute (Couplan, 1998). Flowers of Japanese Honeysuckle (Lonicera japonica) are famous for the little drop of nectar inside and also can be used to adorn salads, cakes and the like. Fruits of at least some honeysuckles mostly from the Pacific Northwest i.e. (L. ciliosa, L. hispidula, L. involucrata and L. villosa) are edible but others are poisonous and unfortunately even the poisonous ones are purported to taste good (Couplan, 1998).
Medicine: Sambucus nigra and S. canadensis are famous medicinals for colds and flu. Japanese Honeysuckle (Lonicera japonica) is a prevalent flu remedy in China used as part of the formulation Yin chiao. Crampbark (Viburnum opulus) and Black Haw (Viburnum prunifolium) have a long history of use for women’s health. Other species have been used medicinally by Native Americans (Moerman, 1998, 2003).
Toxicity: Sambucus contains cyanogenic glycocides and only the stemless fruits should be consumed in any quantity. Some debate exists on the possible toxicity of Red Elderberry (S. racemosa) and a Black Elderberry (S. ebulus) in Europe is considered to be outright poisonous at least in one source (ANWB, 2005). Snowberry fruits (Symphoricarpos spp.) contain saponins and a small amount of chelidonine (Nelson et al., 2007).
Insects: Blackhaw (Viburnum prunifolium) is good for stimulating early brood production. Sambucus nigra is important for pollen. Most Lonicera spp. flowers have tubes too long for bees to reach their nectar. However, some bush and fly honeysuckles (L. morrowi and L. tatarica) are visited by bees. Coralberry (Symphoricarpos orbiculatus) is much sought after by bees when abundant.
Invasiveness: Seven taxa of Honeysuckle (Lonicera spp.) are introduced and invasive to various degrees though a few are not included in Wofford (1989). These include (L. japonica), Breath of Spring (L. fragrantissima), Morrow’s Honeysuckle (L. morrowii) Coral Honeysuckle (L. sempervirens), Standish’s Honeysuckle (L. standishii) Tatarian Honeysuckle (L. tatarica) and Bell’s Honeysuckle (L. x bella) (Miller et al., 2006). Japanese Honeysuckle is by far one of the most widespread exotic invasives covering over 45,000,000 acres according to one analysis (Oswalt, Thompson, & Smith, 2009). The native Honeysuckle (L. sempervirens) makes a great alternative to the Japanese Honeysuckle (Burrell, 2006).
Dye Plants: Elderberries (Sambucus spp.) (Cannon, 1994; Fern, 2009; A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis, 1967).
Other Crafts: The exotic invasive Lonicera japonica can be used for basketry (Hill, 1997).
Aesthetics: Many species are attractive ornamental plants and smell wonderful.
Celastraceae/Bittersweet Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
3 |
Genera |
13 |
Genera |
93 |
Species |
6 |
Species |
35 |
Species |
1200 |
|
|
Lower Taxa |
8 |
|
|
Blue Ridge Genera: Celastrus (2), Euonymus (3), Paxistima (1)
Food: Young parts of Bittersweet (Celastrus spp.) have been consumed in the past after boiling (Couplan, 1998). Not recommended!!!
Toxicity: Celastrus fruits are said to be poisonous containing an unknown gastrointestinal irritant (Nelson et al., 2007). Khat (Catha edulis) is an addictive narcotic used in Africa in places like Somalia (Wink & Van Wyk, 2008).
Insects: Native Bittersweet Celastrus scandens is worked freely by bees.
Invasiveness: The Oriental Bittersweet (Celastrus orbiculatus) is one of the most challenging exotic invasive plants of the Appalachian region. It is distinguished from the native C. scandens that is crosses with by having axillary fruits and different shaped leaves. Winged burning bush (Euonymus alatus) and Winter Creeper (E. fortunei) can be exotic invasives though neither is considered in the Blue Ridge Flora (Miller et al., 2006; Wofford, 1989).
Dye Plants: None known
Other Crafts: Wreathes and baskets can be made from Celastrus orbiculatus though care should be made not to spread invasive fruits or harvest the native which is not common east of the smokies.
Aesthetics: Euonymous spp. and Celastrus spp. have beautiful foliage and fruits in the fall.
Cornaceae/Dogwood Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
1 |
Genera |
1 |
Genera |
7 |
Species |
6 |
Species |
16 |
Species |
105 |
Lower taxa |
2 |
Lower Taxa |
8 |
|
|
Taxonomic commentary: The landscape plant Aucuba japonica has sometimes been included with the Cornaceae. It is now often put in its own family (Aucubaceae) (Heywood et al., 2007). Some debate remains as to whether Nyssaceae is a separate family or a subfamily of Cornaceae (Heywood et al., 2007; Weakley, 2015).
Dogwoods can apparently be broken up into clades. Potentially four sections: Big plants with showy Bracts i.e. C. florida, C. kousa, Little plants with showy bracts i.e. C. canadensis. The biggest group is the small bract group broken into two parts the Cornels i.e. C. mas, C. officinalis, C. sessilis, and a much larger section of which C. alternifolia and C. ammomum are part (“Cornus (genus) - Wikipedia, the free encyclopedia,” n.d.). Edibility seems to follow along these grouping lines.
Blue Ridge Genera: Cornus (1)
Food: Fruits of Bunchberry (C. canadensis) and the introduced landscape plants Cornelian cherry (C. mas) and Asian Dogwood (C. kousa) are edible (Couplan, 1998). The Black Fruited Dogwood (Cornus sessilis) has been enjoyed by friends and myself in California. It is not included in Couplan (1998) but edibility is supported from other sources (Clarke, 1978; Fern, 2008).
Medicine: Cornus canadensis was used medicinally as an anti-malarial during the Civil War (Porcher, 1970).
Toxicity: Most fruits are inedible to people. Bloodtwig dogwood (Cornus sanguinea) can cause skin irritation (Nelson et al., 2007).
Insects: ?
Wildlife: The fruits of Dogwoods are appreciated by many migrating birds and represent an important food source for them.
Invasiveness: None known
Dye Plants: Dogwoods (Cornus spp.) (Fern, 2009; A. Krochmal & C. Krochmal, 1974).
Other Crafts: The wood of Cornus florida is very hard and dense. It may be used for mallets and other durable goods. Dogwood twigs have also been used in basketry and as toothbrushes.
Aesthetics: Many dogwoods are beautiful landscape trees with exceptional bark characteristics. Flowers wilt soon after cutting foliage. The most common native Dogwood Cornus florida is threatened by an anthracnose disease that may remove it from the natural environment.
Cupressaceae/Cypress Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2(n) |
Genera |
7 |
Genera |
30 |
Species |
3(n) |
Species |
34 |
Species |
130 |
Lower taxa |
|
Lower Taxa |
10 |
|
|
Blue Ridge Genera: Juniperus (2), Thuja (1)
Western genera include Incense Cedar (Calocedrus decurrens) which is a major ethnobotanical
of Pacific Northwestern Native Americans (Pojar et al., 1994).
Number for world genera and species from D.J. Mabberly (2008b).
Food: Cones from Junipers (Juniperus spp.) are used in food items such as kraut and distillations i.e. gin, jenever
Toxicity: Can be toxic, use moderately…
Invasiveness: Some species of Juniperus are weedy and have taken over millions of acres (Judd et al., 2008).
Dye Plants: Eastern Red Cedar (Juniperus virginiana) (Eaton, 1973).
Other Crafts: Wood is highly valued and rot resistant.
Aesthetics: Many members of this family are valued in the landscape including Junipers (Juniperus spp.), Psuedo Cypress (Chamaecyparis spp.), Arborvitae (Thuja spp.) and Cypress (Cupressus spp.).
Ericaceae/Heath Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
18(n) |
Genera |
33 |
Genera |
124 |
Species |
50(n) |
Species |
192 |
Species |
4050 |
Lower Taxa |
2 |
Lower Taxa |
112 |
|
|
Taxonomic commentary: This family now includes the formerly distinct Pyrolaceae and Monotropaceae (Heywood et al., 2007). Kartesz (1994) treated these two/three families separately. Their totals are compiled under Ericaceae here. No exotic members are included in Wofford. Heywood et al. include 8 subfamilies and 20 tribes.
Blue Ridge Genera: Arctostaphylos (1), Chamaedaphne (1), Chimaphila (2), Epigaea (1), Gaultheria (1), Gaylussacia (5), Kalmia (2), Leiophyllum (1), Leucothoe (3), Lyonia (2), Menziesia (1), Monotropa (2), Monotropsis (1), Oxydendrum (1), Pieris (1), Pyrola (4), Rhododendron (11), Vaccinium (10)
Food: Fruits from Blueberries/Cranberries (Vaccinium spp.) and Huckleberries (Gaylussacia spp.) are relished for pies, jams, wines etc. Fruits from Manzanita (Arctostphylos spp.) are enjoyed in the western U.S.
Toxicity: Honey from several members of the Ericaceae including Rhododendron spp. may be toxic (Kingsbury, 1964; Nelson, Shih, & Balick, 2007; Pellett, 1977). Mountain laurels (Kalmia spp.), Fetterbush (Pieris spp.) and Dog Hobble (Leucothoe spp.) are all toxic to livestock. Their wood should also probably not be burned. Sodium channel activators are present in Mtn Laurel (Kalmia spp.), Fetterbush (Pieris spp.), Maleberries (Lyonia spp.), Rhododendron spp., Pernettya spp. and Doghobble (Leucothoe spp.) (Nelson et al., 2007). Grayanotoxins or andromedatoxins are usually attributed to Ericaceae toxicity.
Insects: Vaccinium and Gaylussacia provide good surplus nectar for bees. A special honey is produced in Appalachia from the Sourwood (Oxydendrum arboreum). Bees are also fond of Fetterbush (Pieris spp.).
Invasiveness: None of the many species included in the Blue Ridge Flora is introduced (Wofford, 1989)!
Dye Plants:
Heath (Erica spp.) & Heather (Calluna vulgaris) (Cannon, 1994; Fern, 2009)
Madrone (Arbutus menziesii) and Bear berry (Arctostaphylos uva-ursi) (Bliss, 1994; Fern, 2009; A. Krochmal & C. Krochmal, 1974).
Mountain Laurel (Kalmia latifolia) (Eaton, 1973; Fern, 2009; A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis, 1967).
Blueberry (Vaccinium) spp. (Bliss, 1994; Fern, 2009; A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis, 1967)
Other Crafts: Wood from Rhododendron spp. and Mountain laurel (Kalmia spp.) is used for ornamental fence railings.
Aesthetics: Many of the plants in this family are used in the landscape especially for their spring flower displays.
Fabaceae/Bean Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
30(n) 7(i) |
Genera |
158 |
Genera |
727-732 |
Species |
80(n) 27(i) |
Species |
1574 |
Species |
19000-19700 |
|
|
Lower Taxa |
876 |
|
|
This is a very important family. Most members aid in fixing nitrogen in the soil symbiotically with special bacteria that live in their roots. Nitrogen is an essential plant nutrient. Therefore, many members of this family are often used as cover crops. Only a few of the members in the temperate zone are woody shrubs or trees but they are almost all significant in various ways. Redbud (Cercis canadensis) amongst some other members from the family does not fix nitrogen.
Taxonomic commentary: The Fabaceae is the third biggest family in the world. The family is so big and diverse that it is often split into three sub families: Mimosoideae, Caesalpinioideae, Papilionoideae (Heywood et al., 2007).
Woody Blue Ridge Genera: Albizia (1), Cercis (1), Cladrastis (1), Gleditisia (1) Pueraria lobata (1), Robinia (3), Wisteria (2) W. sinensis not included in Wofford!
Food: The Fabaceae is also one of the chief sources of protein for vegetarians. Beans (Phaseolus spp.) are native to Central America (Van Wyk, 2005). Peas (Pisum sativum), Chickpeas (Cicer arientinum) and Lentils (Lens spp.) are all sources of protein for people and animal forage. Red Bud (Cercis spp.) and Black Locust (Robinia pseudoacacia) flowers are choice edibles. Red clover (Trifolium pratense) leaves and flowers are good in tea in moderation.
Medicine: Astragalus (Astragalus membranaceus), Senna (Cassia spp.), Licorice (Glycyrrhiza glabra), Red clover (Trifolium pratense), Kudzu (Pueraria lobata) and Silk tree/Mimosa (Albizzia julibrissin) are just a few of the famous medicinals from this family (Van Wyk & Wink, 2004).
Toxicity: Many members are toxic even possibly deadly and plants from this family should never be sampled without absolute positive identification. Alkaloidal convulsant poison containing genera include Gymnocladus, Sophora, Laburnum and Baptisia (Nelson et al., 2007). Alfalfa (Medicago sativa) and Rattlebox (Sesbania spp.) contain saponins (Kingsbury, 1964). Triterpine and steroidal saponins may occur in the Fabaceae in general (Wink & Van Wyk, 2008). Rattleboxes Sesbania and (Crotalaria) contain pyrrolizidine alkaloids (Nelson et al., 2007). Vicia spp. and Trifolium spp. can cause photosensitivity through their effect on the liver (Kingsbury, 1964). Some people are allergic to members of the Fabaceae including soybeans (Glycine max). Aflotoxin is a toxic fungus that can occur on peanuts (Arachis hypogaea) and other nuts. Robinia contains a toxalbumin (Nelson et al., 2007). Wisteria spp. contains wistarine which is a gylcocide (Nelson et al., 2007). Other potentially toxic members include Locoweeds (Astragalus spp.), Senna (Cassia), Sweet peas (Lathyrus spp.) and Lupines (Lupinus spp.) (Blackwell, 1990).
Insects: The following plants are beloved by bees and other insects: Indigo (Baptisia spp.), Red Bud (Cercis canadensis), Clover (Trifolium spp.), Sweet Clovers (Melilotus spp.), Vetch (Vicia spp.) Alfalfa (Medicago sativa), Black Locust (Robinia pseudoacacia), Alsike Clover (Trifolium hybridum), Bird’s Foot Trefoil (Lotus corniculatus) and False Indigo (Amorpha spp.). Scotch Broom (Cytisus scoparius) also provides pollen for bees.
Invasiveness: Silk tree (Albizia julibrissin), Crown Vetch (Securigera varia syn Coronilla varia), Scotch Broom (Cystisus scoparius), Korean Clover, (Kummerowia stipulaceae), Japanese Clover (K. striata), Shrubby Lespedeza (Lespedeza bicolor), Sericea Lespedeza (L. cuneata), Thunberg’s Lespedeza (L. thunbergii), Bird’s Foot Trefoil (Lotus corniculatus), Black Medick (Medicago lupilina), Yellow Sweetclover (Melilotus officianalis), Kudzu (Pueraria lobata), Sicklepod (Senna obtusifolia), Garden Vetch (Vicia sativa), Japanese Wisteria (Wisteria floribunda), Chinese Wisteria (W. sinensis) (Miller et al., 2006). Introduced species of Trifolium are especially prevalent in many cultivated environments.
Dye Plants: At least thirty seven species of Fabaceae are listed as used for dye (Fern, 2008). Appalachian native and naturalized genera include, Indigo (Baptisia australis) Dyer’s Indigo (B. tinctoria) and Broom (Cystisus scoparius).
Other Crafts: Beans from various members i.e. (Caesalpinia, Crotolaria, Enterlobium) have been used for jewelry. Black Locust wood is very rot resistant and is often used for fence posts and even structurally as part of the foundation for houses. Wisteria and Kudzu can be used for basketry.
Aesthetics: Many members of the Fabaceae have a stunning array of attractive characteristics.
Fagaceae/Beech Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
3(n) |
Genera |
5 |
Genera |
8 |
Species |
23(n) |
Species |
95 |
Species |
620-750 |
|
|
Lower Taxa |
138 |
|
|
Blue Ridge Genera: Castanea (2), Fagus (1), Quercus (20) Wofford excluded: Quercus virginiana.
Western members: Bush Chinkapin (Chrysolepsis sempervirens) Tanbark
Oak (Lithocarpus densiflorus)
Food: The Fagaceae represents an incredible food resource from every genus represented in the Blue Ridge. This family contains the Oaks (Quercus spp.), Chestnuts (Castanea spp.) and Beeches (Fagus spp.) Acorns from White Oaks such as (Quercus alba) and Chestnut Oak (Quercus prinus/montana) contain lower tannin amounts and therefore need less leaching. All Oaks produce edible nuts. Some are more choice than others. A number of mushrooms of which many are edible are also associated with the Oaks (Binion et al., 2008). Native Americans far from being passive forest dwellers of Oak/Chestnut forest often set fires to achieve various goals of which some were related to increasing food availability (Delcourt & Delcourt, 1998; Delcourt et al., 1998).
Toxicity: Tannins are present in many members and are not good to consume in high amounts. American Beech (Fagus grandifolia) contains saponins as well (Kingsbury, 1964).
Insects: Fagus grandifolia and Quercus spp. provide pollen for bees. Bees also get honeydew from Quercus spp. galls. Gypsy moths are a pest threatening Oaks currently. Oaks also provide food for many other insects as well (Tallamy, 2009).
Invasiveness: None known
Dye Plants: European Chestnut (Castanea sp.), Fagus spp., Quercus spp. (citation)
Other Crafts: Endless crafts can be made from the woods of all members of the Fagaceae. Acorns from Quercus spp. offer excellent craft opportunities as well. i especially like the variation in Acorn tops which have served as fairy hats to enterprising folk artists like my friend Sarah Hinds. White Oak baskets are one of the mainstays of Appalachian craft (Law & Taylor, 1991).
Aesthetics: Most members of the Fagaceae are stunningly beautiful landscape trees that often hold on to many of their leaves in the fall. They can get to be quite large and old especially in open grown environments.
Miscellaneous: There is a great publication available online detailing the Oaks of eastern North America (Stein, Binion, & Acciavatti, 2012). Another covers the Oaks of California (Pavlik, Muick, & Johnson, 1993). One covers the whole continent (H. A. Miller & Lamb, 1984). Lastly there is a resource for the Oaks of Central America as well (Kappelle, 2008).
Juglandaceae/Walnut Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2(n) |
Genera |
3 |
Genera |
8 |
Species |
9(n) |
Species |
23 |
Species |
60 |
Lower taxa |
2 |
Lower Taxa |
16 |
|
|
Blue Ridge Genera: Carya (7), Juglans (2) Wofford excluded: Pecan (Carya illionensis).
Food: Many species provide edible nuts including Mockernut Hickory (Carya tomentosa) Pecan (Carya illionensis), Sweet Hickory (C. laciniosa), Shagbark Hickory (C. ovata), Black Walnut (Juglans nigra) and Butternut (J. cinerea). The sap from several genera and species has been consumed by various peoples (Couplan, 1998). English Walnut (Juglans regia) is famous the whole world over.
Medicine: The hulls from the Juglans nigra are used as an anti-parasitic. Following the doctrine of signatures English Walnut is purported to be good for brain health.
Toxicity: Juglone in the root of Juglans nigra is alleleopathic to many plants. Juglans nigra can also cause contact dermatitis (Nelson et al., 2007). Aflotoxin is a toxic fungus that can occur on various nuts.
Insects: Carya spp. are important for bees as a source of pollen and honeydew.
Invasiveness: Juglans nigra is prone to come up strong in open farm fields left uncultivated. Every member of the family represented in Wofford is native to the Blue Ridge.
Dye Plants: All of the above species can be used for dyes. Some are colorfast without mordants.
Other Crafts: Juglans spp. and Carya spp. woods are highly valued for a diversity of crafts including veneers, paneling, carving, utensils etc. Carya spp. are also used for BBQ woodchips.
Aesthetics: Most members make striking landscape trees. Hickory leaves have a beautiful mustard yellow color in the fall.
Aromatics: Leaves and bark of Hickory (Carya spp.) and Walnut (Juglans spp.) are aromatic.
Lauraceae/Bay leaf
Blue Ridge |
|
North America |
|
World |
|
Genera |
2 |
Genera |
13 |
Genera |
50 |
Species |
2(n) |
Species |
40 |
Species |
2500-2750 |
Lower taxa |
|
Lower Taxa |
4 |
|
|
Blue Ridge Genera: Lindera (1), Sassafras (1)
Western Genera: California Bay Laurel (Umbellularia californica)
This is an aromatic, ancient and mostly tropical family
Food: Fruits of Spice Bush (Lindera benzoin) are used like allspice. Twigs and leaves of Spice Bush and Sassafras (Sassafras albidum) are used in tea. Roots of Sassafras were used for a traditional root beer. Bay leaf (Laurus nobilis) is popular as a pot herb. Leaves of California Bay (Umbellularia californica) can be used like Bay leaves though they are stronger in taste. Nuts have also been consumed when prepared properly. Cinnamon (Cinnamomum spp.) is famous as a flavoring spice and also is a potent medicinal. The famous fruit Avocado (Persea spp.) is in this family. Red Bay in the same genus (Persea spp.) grows in Florida and its leaves have been used for flavoring.
Toxicity: Presence of the phytochemical safrole has led to disuse of Sassafras. However its toxicity is less than many commonly consumed substances including beer (Duke & Foster, 1999). The master’s thesis of Kate Cummings (2012) is available for consultation on this topic at greater depth in relation to different forms of preparation and the resulting safrole content. Consumption in moderation is probably prudent. California Bay is the carrier of Sudden Oak Death.
Birds: Fruits of various members from the Lauraceae are relished by birds in their long migration south for the winter.
Invasiveness: None known
Dye Plants: Sassafras can be used as a dye plant (Fern, 2008; Krochmal & Krochmal, 1974; Nicholson & Clovis, 1967).
Aesthetics: Most members of the Lauraceae are attractive at all times of year.
Aromatics: Bark, leaves, flowers are often aromatic for most members of the Lauraceae.
Magnoliaceae/Magnolia Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2 |
Genera |
2 |
Genera |
2-3-10 |
Species |
6 |
Species |
11 |
Species |
220 |
Lower taxa |
|
Lower Taxa |
|
|
|
Taxonomic commentary: This is a small family with primitive flower characteristics and a distinct link to Eastern Asia.
Blue Ridge Genera: Liriodendron (1), Magnolia (5)
Food: The copious nectar from Tulip Tree flowers can be consumed. The leaves of Sweet Bay (Magnolia virginiana) are used like bay laurel leaves in cooking. The flowers of Cucumber Magnolia can be eaten and the flowers and buds of several Asian species as well as young and tender leaves have been eaten in different parts of Asia (Couplan, 1998). My friend, colleague and fellow forager Robin Harford makes particularly delectable food items including a syrup from M. liliiflora.
Toxicity: Magnolia grandiflora can cause contact dermatitis (Nelson et al., 2007).
Insects: Tulip tree (L. tulipifera) is one of the major bee trees in the Appalachian region.
Invasiveness: None known though Tulip Tree can quickly take over abandoned farm fields and outpace the growth of sometimes more desirable hardwoods in forest situations due to a strategy of long lasting buried seeds.
Dye Plants: Liriodendron tulipifera can be used as a dye (Fern, 2009; A. Krochmal & C. Krochmal, 1974).
Aesthetics: The exotic but not invasive Saucer Magnolia (M. x soulangeana) and Star Magnolia (M. stellata) make beautiful landscape plants. Many species native to north America are planted for ornament as well.
Aromatics: Many members of the Magnoliaceae have fabulous smells.
Other Crafts: The fruiting structure of Magnolia can be used ornamentally. Magnolia wood is a classic one used in carving and making bowls. Tulip tree wood is also used in carving, basket making and wood siding.
Malvaceae/Mallow Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
3(n) 2(i) |
Genera |
41 |
Genera |
115 |
Species |
5(n) 8(i) |
Species |
260 |
Species |
2000 |
Lower taxa |
2 |
Lower Taxa |
93 |
|
|
Blue Ridge Genera: Abutilon (1), Hibiscus (3,2), Iliamna (1), Malva (3), Sida (1,2)
Wofford excluded Althea rosea, Malva rotundifolia and Modiola caroliniana.
Food: Okra (Abelmoschus esculentus) is a cultivated crop that is famous in the south. Most leaves can be consumed raw, cooked or as tea including Marshmallow (Althaea officinalis), Mallow (Malva spp.) flowers are edible as are almost all species in the family. The calyces of Cranberry Hibiscus or Roselle (Hibiscus acetosa and H. sabdariffa) make a fabulous beverage. Rose of Sharon (Hibiscus syriacus) is imminently edible for flowers and cooked greens. Chocolate (Theobroma cacao) formerly from the Sterculiaceae is now in this family as well.
Toxicity: The Malvaceae is generally considered a non-toxic family with the exception of cottons (Gossypium spp.)
Insects: Hollyhock (Althaea rosea) is popular with bees.
Invasiveness: Rose of Sharon (Hibiscus syriacus) can become locally invasive. Malva spp. can spread readily.
Dye Plants: Cotton (Gossypium spp.) (A. Krochmal & C. Krochmal, 1974). Hibiscus spp. (Buchanan, 1995; Fern, 2009) and Malva spp. (Bliss, 1994; Fern, 2009).
Aesthetics: Many members have beautiful flowers including Hibiscuses (Hibiscus spp.) and Hollyhocks (Alcea spp.).
Pinaceae/Pine Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
4(n) |
Genera |
7 |
Genera |
12 |
Species |
11(n) |
Species |
72 |
Species |
225 |
Lower taxa |
|
Lower Taxa |
35 |
|
|
Blue Ridge Genera: Abies (2), Picea (1), Pinus (6), Tsuga (2)
Wofford excluded Larch (Larix decidua) and Norway spruce (Picea abies)
Number for world genera and species from D.J. Mabberly (2008b).
Food: Tea and beer can be made from the needles of Pine (Pinus spp.) and Spruces (Picea spp.). The sap from Picea spp. can be used like a chewing gum according to a personal conversation with Doug Elliot.
Toxicity: The smoke from resinous yellow Pines especially their cones should not be inhaled in large quantities. Balsam Fir (Abies balsamea) can cause contact dermatitis (L. S. Nelson, Shih, & Balick, 2007). The resin present in needles should also not be consumed in large quantities and a cold versus hot extraction in tea is preferred for this reason (Elpel, 2013).
Insects: Pine Bark beetles are a problem from coast to coast in the United States.
Dye Plants:
Spruce (Picea spp.) (A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis,
1967)
Hemlocks (Tsuga canadensis and T. caroliniana) (Fern, 2009; A. Krochmal & C. Krochmal,
1974; Moerman, 1998; Nicholson & Clovis, 1967)
Pines (Pinus spp.) (Eaton, 1973; Fern, 2008)
Other Crafts: Endless crafts can be pursued using the cones alone. White pine wood is generally considered more choice than yellow pine wood. White pines tend to have needles grouped in 5’s. This is probably the most important family in the world for timber.
Aesthetics: Many species make great evergreen landscape trees and are often used as Christmas trees as well. Virginia Pine (Pinus virginiana) can tend to look trashy as it gets older with many dead persistent branches and Eastern White Pine (P. strobus) is susceptible to tremendous damage even in large healthy trees during winter storms. That said many of the most stunning trees of the hundreds of species i have met are in the pine family mostly from the western United States including Bristlecone Pine (P. aristata) Sugar Pine (P. lambertiana), Coulter’s Pine (P. coulteri), Gray Pine (P. sabiana) Ponderosa Pine (P. ponderosa) However, i have a special place in my heart for the southeastern Longleaf Pine (P. palustris) who’s ecology, ethnobotany, tragic history and potential restoration has been documented extensively (Earley, 2006; Finch, 2012; Jose et al., 2007; Latham, 2013; Neel, Sutter, Way, & Franklin, 2012; Outland, 2004; Streich, 2012; Way, 2011).
Rosaceae/Rose Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
20(n) 3(i) |
Genera |
67 |
Genera |
85-100 |
Species |
83(n) 19(i) |
Species |
842 |
Species |
2000 |
Lower taxa |
|
Lower Taxa |
406 |
|
1,300-1500 |
Taxonomic commentary: The taxonomy of Hawthorns (Crataegus spp.) and Juneberries (Amelanchier spp.) is very messy (Lance, 2004). A whole book has been dedicated to the Hawthorns alone in an attempt to whittle down the over 1,000 named species in the southeast to around 200 proper species after eliminating synonyms (Lance, 2014).
Blue Ridge Genera: Agrimonia (5), Alchemilla (1), Amelanchier (5), Aronia (2), Aruncus (1) Crataegus (19,1), Dalibarda (1), Duchesnea (1), Filipendula (1), Fragaria (2), Geum (7), Malus (2,1), Physocarpus (1), Porteranthus (2), Potentilla (4,2), Prunus (7,4), Pyrus (1), Rosa (5,4), Rubus (11,3), Sanguisorba (1), Sorbus (1), Spiraea (5,1), Waldsteinia (2)
Wofford excluded: Potentilla anserine, P. intermedia, P. rivalis, Prunus caroliniana, Rosa bracteata, R. damascena, R. gallica, Rubus discolor, R. illecebrosus, Sanguisorba minor, Spiraea prunifolium, S. salicifolia, S. x vanhouttei.
Food: This is one of the major families for fruits in the temperate world including Apples (Malus spp.), Pears (Pyrus spp.), Quince (Cydonia, Chaenomeles) Strawberry (Fragaria spp.). Almonds, Apricots, Cherries, Nectarines, Peaches, and Plums are all in the same genus (Prunus spp.). Juneberries (Amelanchier spp.), Hawthorns (Crataegus spp.), Chokeberry (Aronia spp.) and Mountain Ash/Rowan (Sorbus spp.) are a few wild genera that are foraged for food. Many shrubs including cane fruits (Rubus spp.) and Roses (Rosa spp.) also are edible. Apple wood is used to smoke food as well.
Toxicity: Cyanogenic glycocides are present in some members of the Rosaceae (L.S. Nelson et al., 2007). Some people are allergic to members of the Rosaceae such as Strawberries. Agrimony (Agrimonia spp.) and Tea Rose (Rosa odorata) can cause contact dermatitis in particular (Nelson et al., 2007). Triterpine saponins occur in the Rosaceae too (Wink & Van Wyk, 2008).
Insects: Most plants in the Rosaceae provide copious amounts of early pollen and nectar for bees. Japanese Beetles can be a pest on members of this family. Black Cherry (Prunus serotina) is one of the greatest supporters/targets of insects (Tallamy, 2009).
Invasiveness: Shrubby Blackberry (Rubus fruticosus) and Eelkek (R. mollucana) both are not listed in Wofford (1989) but are high priority invasives for many states. Wineberry (R. phoenicolasius) can be somewhat invasive locally. Wild Strawberry (Duchesnea indica), Sulphur Cinquefoil (Potentilla recta) and Bradford pear (Pyrus calleryana) can be invasive. Japanese spiraea (Spiraea japonica) is an exotic invasive shrub. Multiflora rose (Rosa multiflora) is one of the worst exotic invasives in the eastern U.S. covering many million acres alone (Miller et al., 2006; Wofford, 1989) However, some predators both microbiological and entomological are working to reduce the multiflora population (Amerine, 2000; Kay, Lewis, & Langeland, 1995).
Dye Plants: Apples (Malus spp.) Cinquefoil (Potentilla spp.), Pear (Pyrus communis), Plum (Prunus americana), Peach (P. persica), Wild Black Cherry (P. serotina). Brambles (Rubus fruticosus, R. idaeus, and R. tricolor). An incredible 124 species of Prunus and 158 species of Rubus have been listed as dyes (Fern, 2008)!
Other Crafts: Wood is good for carving.
Aesthetics: Many members of the Rosaceae have beautiful flowers, attractive fruits and nice bark.
Salicaceae/Willow Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
2 |
Genera |
2 |
Genera |
2 |
Species |
10(n) 4(i) |
Species |
117 |
Species |
385 |
Lower taxa |
3 |
Lower Taxa |
95 |
|
|
Blue Ridge Genera: Populus (3,2) Salix (7,2)
Wofford excluded: Populus x canescens, Populus nigra, Salix caprea, S. cinerea ssp. cinerea, S. cinerea ssp. oleifolia, S. pentandra, and S. purpurea.
Medicine: Willows are famous for their salicylic acid compounds that act like aspirin to bring about pain relief when applied in sufficient dosage (Foster & Hobbs, 2002; Van Wyk & Wink, 2004).
Toxicity: ?
Insects: Willows (Salix spp.) provide early food to bees in the late winter time.
Invasiveness: Silver poplar (Populus alba) can be invasive (Miller et al., 2006).
Dye Plants: Many willows have been used as dyes including (S. alba, S. caprea, S. cinerea, S. fragilis, S. purpurea, and S. viminalis)
Other Crafts: Many species of Willows are used for basketry (Trestain, 1998). Living fences and structures have also been formed out of Willow. Various species are coppiced. The unopened inflorescences are famous as additions to floral arrangements as well.
Aesthetics: Many species of both genera are valued landscape plants especially Weeping Willow (Salix babylonica). Willows and Poplars are some of the most cold hardy deciduous trees. They light up places with their beautiful yellow leaves in the fall.
Vitaceae/Grape Family
Blue Ridge |
|
North America |
|
World |
|
Genera |
3(n) |
Genera |
4 |
Genera |
12-14 |
Species |
11(n) |
Species |
40 |
Species |
800 |
Lower taxa |
|
Lower Taxa |
9 |
|
|
Blue Ridge Genera: Ampelopsis (2), Parthenocissus (1), Vitis (8) Wofford excluded: Ampelopsis brevipendunculata
Food: Grapes (Vitis spp.) are commonly used fresh and for jams, jellies, juices, wines, etc. Some are bigger and better than others. Leaves of Vitis spp. have also been consumed most famously as dolmas from Greece.
Toxicity: Virginia Creeper (Parthenocissus quinquefolia) and Boston Ivy (P. tricuspidata) contain irritant raphides (Nelson et al., 2007). Ampelopsis spp?
Insects: Ampelopsis spp. and Parthenocissus spp. are good honey plants.
Invasiveness: Ampelopsis aconitifolia?
Dye Plants:
Grapes (Vitis spp.) (Fern, 2009; A. Krochmal & C. Krochmal, 1974; Nicholson & Clovis, 1967).
Virginia creeper (Parthenocissus quinquefolia) (Fern, 2008).
Other Crafts: The vines of various species can be used to make baskets but should only be used when abundant.
For the next class we will cover major Medicinal families and it will be posted around October 28th
My apologies if you have found yourself waiting for this last class to be posted…Even after 6 years of teaching this class and several more years researching materials for it i sometimes find it hard to carve out the time to edit and update as much as i would like. This year has been particularly hard in that regard with blocks of computer time few and far between with so much work out in the field…i had hoped to recruit some help in other arenas through apprentices but a lot of that has fallen through. Remember that past classes from the last year are up and available if you simply can’t wait for new versions to come out….
Below are items to think about/comment on. Please write me directly at marc@botanyeveryday.com or leave information at the Facebook group.
i WOULD REALLY LOVE TO HEAR WHAT YOU HAVE TO SAY!!!
- Make a list of the woody plants around you and share that info with some people
-Start a collection of leaves from the various species you can find
- Think of a craft you can make with woody plants
- Attend a workshop or a class and write up a brief description of plants or information learned.
- Post any clear photos of question plants to Facebook or send in an email.
Praises to all that have donated to the cause. i encourage everyone reading this to donate as they are able financially, commentarialy, or energetically... Your contributions greatly help me continue this crucial work of ethnobotanical research and education. Please let me know your thoughts in general and anyway i can help this class serve you best.
Thanks, marc
References
Abrams, M. D., & Nowacki, G. J. (2008). Native Americans as Active and Passive Promoters of Mast and Fruit Trees in the eastern USA. The Holocene, 18(7), 1123–1137. https://doi.org/10.1177/0959683608095581
Altman, N. (2000). Sacred trees: spirituality, wisdom & well-being. New York: Sterling Pub.
Amerine, J. W. (2000, November). Biological Control Agents of Multiflora Rose. West Viriginia Cooperative Extension. Retrieved from http://www.wvu.edu/~agexten/ipm/weeds/multiflor.htm
ANWB. (2005). Bomen van Europa. Den Haag, The Netherlands: ANWB.
Baldwin, B. G., Goldman, D. H., Keil, D. J., Patterson, R., & Rosatti, T. J. (Eds.). (2012). The Jepson Manual: Vascular Plants of California (2nd ed.). Berkeley, CA: University of California Press.
Bash, B. (2002). Ancient Ones: The World of the Old-Growth Douglas Fir. San Francisco: Sierra Club Books For Children.
Beattie, M., & Levine, L. (1993). Working with Your Woodland: A Landowner’s Guide (Revised Ed). Hanover, NH: University Press of New England.
Binion, D., Stephenson, S., Roody, W., Burdsall, H. H., Miller, O. K., & Vasilyeva, L. (2008). MACROFUNGI ASSOCIATED WITH OAKS OF EASTERN NORTH AMERICA. Morgantown, WV: West Virginia University Press.
Black, B. A., Ruffner, C. M., & Abrams, M. D. (2006). Native American influences on the forest composition of the Allegheny Plateau, northwest Pennsylvania. Canadian Journal of Forest Research, 36(5), 1266–1275. https://doi.org/10.1139/x06-027
Bliss, A. (1993). North American Dye Plants. Loveland, CO: Interweave Press.
Bowers, J. E. (1998). Flowers and Shrubs of the Mojave Desert (First Edition, 1st Printing). Western Natl Parks Assoc.
Bowers, J., & Wignall, B. (1993). Shrubs & Trees of the Southwest Deserts. Western Natl Parks Assoc.
Brockman, F. (1986). Trees of North America (Rev ed). New York: Golden Press.
Brosi, S. (2010, August). Steps Toward Butternut (Juglans cinerea L.) Restoration (Dissertation). University of Tennessee, Knoxville, TN. Retrieved from http://trace.tennessee.edu/utk_graddiss/779
Bryant, G. (1996). Growing Rhododendrons and Azaleas. London : New York, NY: Cassell.
Bryant, G. (2001). Rhododendrons and Azaleas. Willowdale, Ont. ; Buffalo, N.Y: Firefly Books.
Buffin, M., W. (2005). Winter-flowering Shrubs. Portland, OR: Timber Press.
Burrell, C. C. (2006). Native Alternatives to Invasive Plants. Brooklyn, NY: Brooklyn Botanic Garden.
Caldecott, M. (1993). Myths of the Sacred Tree (Original). Destiny Books.
Cannon, J., & Cannon, M. (2003). Dye Plants and Dyeing. Portland, OR: Timber Press.
Carter, J. L. (1997). Trees and Shrubs of New Mexico. Jack L Carter.
Castner, J. (2004). Photographic Atlas of Botany and Guide to Plant Identification. Gainesville, FL: Feline Press.
Céspedes, F. C., & Lindquist, E. S. (2007). Árboles Comunes de la Reserva Natural Absoluta Cabo Blanco Costa Rica / Common Trees of Cabo Blanco Absolute Nature Reserve. INBio Editorial.
Clark, S. L., Brosi, S. L., Schlarbaum, S. E., & Grissino-Mayer, H. D. (2008). Dendrochronology of two butternut (Juglans cinerea) populations in the southeastern United States. Forest Ecology and Management, 255(5–6), 1772–1780. https://doi.org/10.1016/j.foreco.2007.11.040
Clarke, C. B. (1978). Edible and Useful Plants of California. Berkeley, CA: University of California Press.
Condit, R., Pérez, R., & Daguerre, N. (2010). Trees of Panama and Costa Rica. Princeton University Press.
Corbett, J., & Corbett, M. (2000). Designing Sustainable Communities: Learning from Village Homes. Washington, D.C.: Island Press.
Core, E. L., & Ammons, N. (1999). Woody plants in winter a manual of common trees and shrubs in winter in the northeastern United States and southeastern Canada. Morgantown, WV: West Virginia University Press.
Cornus (genus) - Wikipedia, the free encyclopedia. (n.d.). Retrieved September 24, 2011, from http://en.wikipedia.org/wiki/Cornus_(genus)
Couplan, F. (1998). The Encyclopedia of Edible Plants of North America. New Canaan, CT: Keats Pub.
Cox, K. (2005). Rhododendrons & Azaleas: A Colour Guide. Ramsbury: Crowood Press.
Cox, K. N. E., & Glen, L. (1998). Rhododendrons. San Diego, CA: Thunder Bay Pr.
Cox, P. A., & Cox, K. N. (1988). Encyclopedia of Rhododendron Hybrids. Portland, OR: Timber Press, Incorporated.
Coyier, D. L., & Roane, M. K. (1986). Compendium of Rhododendron and Azalea Diseases (illustrated edition edition). St. Paul, Minn., USA: Amer Phytopathological Society.
Crawford, M. (2010). Creating a Forest Garden: Working with Nature to Grow Edible Crops. Green Books.
Creasy, R. (1982). The Complete Book of Edible Landscaping: Home Landscaping with Food-Bearing Plants and Resource-Saving Techniques. San Francisco: Sierra Club Books.
Cullen, J. (2005). Hardy Rhododendron Species: A Guide to Identification. Portland, OR: Timber Press, Incorporated.
Cummings, K. (2012, May). Sassafras Tea: Using a Traditional Method of Preparation to Reduce the Carcinogenic Compound Safrole (Master’s Thesis for an M.S. in Forestry). Clemson University, Clemson, SC.
Curtis, M. A. (1945). The Shrubs and Woody Vines of North Carolina. Raleigh: Dept. of Conservation and Development.
Dalgleish, H. J., & Swihart, R. K. (2012). American Chestnut Past and Future: Implications of Restoration for Resource Pulses and Consumer Populations of Eastern U.S. Forests. Restoration Ecology, 20(4), 490–497. https://doi.org/10.1111/j.1526-100X.2011.00795.x
Davidian, H. H. (1982). The Rhododendron Species, Vol. 1: The Lepidotes (1st Ed). Portland, OR: Timber Press, Incorporated.
Davidian, H. H. (2003a). The Rhododendron Species: Azaleas. Portland, OR: Timber Press, Incorporated.
Davidian, H. H. (2003b). The Rhododendron Species: Elepidotes. Portland, Ore: Timber Press, Incorporated.
Davidian, H. H. (2003c). The Rhododendron Species: Elepidotes Continued (Neriiflorum - Thomsonii, Azaleastrum and Camtschaticum). Portland, Or: Timber Press, Incorporated.
Delcourt, P. A., & Delcourt, H. R. (1998). The Influence of Prehistoric Human-Set Fires on Oak-Chestnut Forests in the Southern Appalachians. Castanea, 63(3), 337–345. https://doi.org/10.2307/4033982
Delcourt, P. A., Delcourt, H. R., Ison, C. R., Sharp, W. E., & Gremillion, K. J. (1998). Prehistoric Human Use of Fire, the Eastern Agricultural Complex, and Appalachian Oak-Chestnut Forests: Paleoecology of Cliff Palace Pond, Kentucky. American Antiquity, 63(2), 263–278. https://doi.org/10.2307/2694697
Devall, B. (1993). Clearcut: The Tragedy of Industrial Forestry. Japan: Sierra Club Books, Earth Island Press.
Dirr, M. A. (1997). Dirr’s Hardy Trees and Shrubs an Illustrated Encyclopedia. Portland, OR: Timber Press.
Dirr, M. A. (1998). Manual of Woody Landscape Plants (5th ed). Athens, GA: Stipe Publishing LLC.
Dreistadt, S. H. (2004). Pests of Landscape Trees and Shrubs: An Integrated Pest Management Guide (2nd Ed). Oakland, CA: Univ of California Agriculture & Natural Resources.
Duke, J. A., & Foster, S. (1999). A Field Guide to Medicinal Plants and Herbs: Of Eastern and Central North America (Peterson Field Guides) (1st ed.). New York: Houghton Mifflin.
Duke, J. A., & Foster, S. (2014). Peterson Field Guide to Medicinal Plants and Herbs of Eastern and Central North America (3rd Ed). Houghton Mifflin Harcourt.
Dwyer, J. F., Nowak, D. J., Noble, M. H., & Sisinni, S. M. ; (2000). Connecting people with ecosystems in the 21st century: an assessment of our nation’s urban forests. Retrieved from http://www.treesearch.fs.fed.us/pubs/29448
Earle, C. J., Spellenberg, R., & Nelson, G. (2014). Trees of Western North America. (A. Hughes K., Ed.). Princeton, NJ: Princeton University Press.
Earley, L. S. (2006). Looking for Longleaf: The Fall and Rise of an American Forest (Reprint Ed). Chapel Hill: The University of North Carolina Press.
Eaton, A. H. (1973). Handicrafts of the Southern Highlands. New York: Dover Publications.
Eckenwalder, J. E. (2009). Conifers of the world: the complete reference. Portland: Timber Press.
Edwards, A. L., & Bennett, B. C. (2005). Diversity of Methylxanthine Content in Ilex cassine L. and Ilex vomitoria Ait.: Assessing Sources of the North American Stimulant Cassina. Economic Botany, 59(3), 275–285. https://doi.org/10.1663/0013-0001(2005)059[0275:DOMCII]2.0.CO;2
Elias, T. (1980). The Complete Trees of North America, Field Guide and Natural History. New York: Van Nostrand Reinhold.
Elmore, F. H. (1976). Shrubs and Trees of the Southwest Uplands. Western Natl Parks Assoc.
Elpel, T. (2013). Botany in a Day: The Patterns Method of Plant Identification. Pony, MT: HOPS Press, LLC.
Escobedo, F. J., & Nowak, D. J. (2009). Spatial heterogeneity and air pollution removal by an urban forest. Landscape and Urban Planning, 90(3–4), 102–110. https://doi.org/10.1016/j.landurbplan.2008.10.021
Evert, R. F., Eichhorn, S. E., & Raven, P. H. (2012). Biology of Plants (8th Ed). New York: W. H. Freeman.
Farjon, A. (2008). A natural history of conifers. Portland, Or.: Timber Press.
Farjon, A. (2010). A handbook of the world’s conifers. Leiden; Boston: Brill.
Fern, K. (2008). Plants for a future - 7300 useful plants database. Retrieved April 23, 2009, from http://www.pfaf.org/index.php
Finch, B. (2012). Longleaf, Far as the Eye Can See: A New Vision of North America’s Richest Forest. Chapel Hill, NC: The University of North Carolina Press.
Flint, H. L. (1997). Landscape Plants for Eastern North America: Exclusive of Florida and the Immediate Gulf Coast (2nd Ed). New York: Wiley.
Foote, L. E., & Jones, S. B. (1989). Native Shrubs and Woody Vines of the Southeast: Landscaping Uses and Identification. Portland, OR: Timber Press.
Ford, A., & Nigh, R. (2015). Maya Forest Garden: Eight Millennia of Sustainable Cultivation of the Tropical Woodlands. Left Coast Press.
Gilkey, H. M., & Packard, P. (2001). Winter twigs: a wintertime key to deciduous trees and shrubs of northwestern Oregon and western Washington. Corvallis, OR: Oregon State University Press.
Gilman, E. (2001). Illustrated Guide to Pruning (2nd Ed.). Albany, NY: Cengage Learning.
Gilman, E. F. (1997). Trees for Urban and Suburban Landscapes. Albany: Cengage Learning.
Gregory McPherson, E. (1992). Accounting for benefits and costs of urban greenspace. Landscape and Urban Planning, 22(1), 41–51. https://doi.org/10.1016/0169-2046(92)90006-L
Grimm, W. C. (1966). Recognizing Native Shrubs. Harrisburg, PA: Stackpole Books.
Group, T. A. P. (2009). An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III. Botanical Journal of the Linnean Society, 161(2), 105–121. https://doi.org/10.1111/j.1095-8339.2009.00996.x
Hageneder, F. (2005). The meaning of trees: botany, history, healing, lore. San Francisco: Chronicle Books.
Hansen, A. L., Severson, M., & Waterman, D. L. (2011). A Landowner’s Guide to Managing Your Woods: How to Maintain a Small Acreage for Long-Term Health, Biodiversity, and High-Quality Timber Production. North Adams, MA: Storey Publishing, LLC.
Hardin, J. W., Leopold, D. J., & White, F. M. (2001). Harlow and Harrar’s Textbook of Dendrology (9th ed.). New York: McGraw-Hill.
Harlow, W. M. (1946). Fruit key and twig key to trees and shrubs: fruit key to northeastern trees, twig key to the deciduous woody plants of eastern North America. New York: Dover Publications.
Hart, R. A. de J. (Robert A. de J. (1996). Forest Gardening: Cultivating an Edible Landscape. White River Junction, VT: Chelsea Green Pub. Co.
Hawthorne, L. (2014). Rhododendrons: An illustrated guide to varieties, cultivation and care, with step-by-step instructions and over 135 beautiful photographs (Ill edition). S.l.: Southwater.
Hepting, G. H. (1971). Diseases of Forest and Shade Trees of the United States. USDA Forest Service.
Heywood, V. H., Brummitt, R. K., Culham, A., & Seberg, O. (2007). Flowering Plant Families of the World (Revised). Buffalo, NY; Richmond Hill, Ont.: Firefly Books.
Hickman, J. C. (1993). The Jepson Manual: Higher Plants of California. Berkeley, CA: University of California Press.
Hill, S. H. (1997). Weaving New Worlds: Southeastern Cherokee Women and Their Basketry. Chapel Hill, NC: University of North Carolina Press.
Hilts, S., & Mitchell, P. (2009). The Woodlot Management Handbook: Making the Most of Your Wooded Property For Conservation, Income or Both (2nd Ed, Revised & Expanded). Richmond Hill, Ont.: Firefly Books.
Houtman, R. (2004). Variegated Tress and Shrubs: The Illustrated Encyclopedia. Portland, OR: Timber Press.
Huegel, C. N. (1995). Florida plants for wildlife: A selection guide to native trees and shrubs. Orlando, FL: Florida Native Plant Society.
Hugo, N. R. (2011). Seeing Trees: Discover the Extraordinary Secrets of Everyday Trees. Timber Press.
Hyams, E. (1965). Ornamental Shrubs for Temperate Zone Gardens. New York: A.S. Barnes & Co.,, South Brunswick:
Jacke, D., & Toensmeier, E. (2005a). Edible Forest Gardens: Ecological Design and Practice for Temperate Climate Permaculture (Vol. 2). White River Junction, VT: Chelsea Green Pub. Co.
Jacke, D., & Toensmeier, E. (2005b). Edible Forest Gardens: Ecological Vision and Theory for Temperate Climate Permaculture (Vol. 1). White River Junction, VT: Chelsea Green.
Jensen, D., & Draffan, G. (2003). Strangely Like War. White River Junction, VT: Chelsea Green Pub. Co.
Jiménez Madrigal, Q. (2002). Arboles maderables de Costa Rica: ecología y silvicultura = Timber trees of Costa Rica : ecology and silviculture. Santo Domingo, Heredia, Costa Rica: InBio, Instituto Nacional de Biodiversidad.
Jones, R. L., & Wofford, B. E. (2013). Woody Plants of Kentucky and Tennessee: The Complete Winter Guide to Their Identification and Use. Lexington, KY: The University Press of Kentucky.
Jonnes, J. (2016). Urban Forests: A Natural History of Trees and People in the American Cityscape. New York: Viking.
Jose, S., Jokela, E. J., & Miller, D. L. (2007). The Longleaf Pine Ecosystem: Ecology, Silviculture, and Restoration. New York: Springer.
Judd, M. (2013). Edible Landscaping with a Permaculture Twist: How to Have Your Yard and Eat It Too. Ecologia.
Judd, W. S., Campbell, C. S., Kellog, E. A., Stevens, P. F., & Donahue, M. J. (2008). Plant Systematics: A Phylogenetic Approach (3rd ed.). Sunderland, MA: Sinauer Associates.
Kappelle, M. (2008). Biodiversidad de los Bosques de Roble (encino) de la América tropical / Biodiversity of the Oak Forests of Tropical America. Heredia, Costa Rica: Editorial INBio.
Kartesz, J. T. (1994). A Synonymized Checklist of the Vascular Flora of the United States, Canada, and Greenland (2nd ed). Portland, OR: Timber Press.
Kay, S. H., Lewis, W. M., & Langeland, K. A. (1995). Integrated Management of Multiflora Rose in North Carolina. North Carolina Cooperative Extension.
Keeler, H. L. (1969). Our northern shrubs and how to identify them. New York: Dover Publications.
Kelly, D., & Braasch, G. (1988). Secrets of the Old Growth Forest. Salt Lake City, UT: Peregrine Smith Books.
Kingsbury, J. M. (1964). Poisonous Plants of the United States and Canada. Englewood Cliffs, NJ: Prentice-Hall.
Kourik, R., & Kane, M. (1986). Designing and Maintaining Your Edible Landscape Naturally. Santa Rosa, CA : Metamorphic Press ; Emmaus, PA : Distribution, Rodale Press.
Krochmal, A., & Krochmal, C. (1974). The Complete Illustrated Book of Dyes from Natural Sources. Garden City, NY: Doubleday.
Krochmal, A., & Krochmal, C. (1982). Uncultivated Nuts of the United States. Washington, D.C.? U.S. Dept. of Agriculture, Forest Service.
Kurz, H., & Godfrey, R. (1962). Trees of Northern Florida. Gainesville, FL: University Press of Florida.
Lance, R. (2004). Woody Plants of the Southeastern United States: A Winterguide. Athens, GA: University of Georgia Press.
Lance, R. (2014). Haws; A Guide to Hawthorns of the Southeastern United States. Mills River, NC: Ron Lance.
Lansky, M. (1992). Beyond the Beauty Strip; Saving What’s Left of Our Forests. Gardiner, ME: Tilbury House Publishers.
Latham, D. (2013). Painting the Landscape with Fire: Longleaf Pines and Fire Ecology. Columbia, SC: University of South Carolina Press.
Law, R. N., & Taylor, C. W. (1991). Appalachian white oak basketmaking: handing down the basket. Knoxville: University of Tennessee Press.
Leach, D. G. (1961). Rhododendrons of the World (1ST edition). New York: Scribner.
Leiva, A. (2007). Trees of Cuba. Oxford: Macmillan Caribbean.
Little, C. E. (1995). The dying of the trees: The pandemic in America’s forests. New York, NY: Viking.
Little, E. L. (1976). Atlas of United States Trees Volume 3: Minor Western Hardwoods. Washington, D.C.: United States Government Printing Office.
Little, E. L. (1978). Atlas of United States Trees Volume 5. Florida. United States Department of Agriculture Forest Service.
Little, E. L. (n.d.). Atlas of United States Trees; Volume 1. Conifers and Important Hardwoods (First Edition edition). U.S. Department of Agriculture.
Little, E. L., & Wadsworth, F., H. (1964). Common Trees of Puerto Rico and the Virgin Islands; Agriculture Handbook No. 249 (Ag Handbook No. 249 edition). Washington, DC: U. S. Department of Agriculture Forest Service.
Lovell, H. B. (1977). Honey Plants. (L. R. Goltz, Ed.) (Rev ed.). Medina, OH: Gleanings in Bee Culture.
Lovell, J. H. (1999). Honey Plants of North America. A I Root Co.
Mabberley, D. J. (2008a). Mabberley’s plant-book: a portable dictionary of plants, their classification and uses. Cambridge, UK; New York: Cambridge University Press.
Mabberley, D. J. (2008b). Mabberley’s Plant-Book: A Portable Dictionary of Plants, Their Classification and Uses. Cambridge, UK ; New York: Cambridge University Press.
MacKay, P. (2003). Mojave Desert Wildflowers: A Field Guide to Wildflowers, Trees, and Shrubs of the Mojave Desert, Including the Mojave National Preserve, Death Valley National Park, and Joshua Tree National Park (1st ed.). FalconGuides.
Maloof, J. (2011). Among the Ancients: Adventures in the Eastern Old-Growth Forests. Ruka Press.
Marcelle, G., Hawthorn, W., & Jules, D. (2004). Caribbean Spice Island Plants: Trees, Shrubs and Climbers of Grenada, Carriacou and Petit Martinique: a Picture Gallery with Notes on Identification, Historical and Other Trivia. Oxford: Oxford Forestry Institute.
Marshall, R. C. (1939). Silviculture of the Trees of Trinidad and Tobago, British West Indies. Oxford University Press.
Marshall, R. C., Trinidad, & Forest Department. (1930). Notes on the silviculture of the more important timber trees of Trinidad and Tobago, with information on the formation of woods. Trinidad: Govt. Print. Off.
McEwan, R. W., Dyer, J. M., & Pederson, N. (2011). Multiple interacting ecosystem drivers: toward an encompassing hypothesis of oak forest dynamics across eastern North America. Ecography, 34(2), 244–256. https://doi.org/10.1111/j.1600-0587.2010.06390.x
McHale, M. R., Gregory McPherson, E., & Burke, I. C. (2007). The potential of urban tree plantings to be cost effective in carbon credit markets. Urban Forestry & Urban Greening, 6(1), 49–60. https://doi.org/10.1016/j.ufug.2007.01.001
McPherson, E. G., Nowak, D., Heisler, G., Grimmond, S., Souch, C., Grant, R., & Rowntree, R. (n.d.). Quantifying urban forest structure, function, and value: the Chicago Urban Forest Climate Project. Urban Ecosystems, 1(1), 49–61. https://doi.org/10.1023/A:1014350822458
McPherson, E. G., & Simpson, J. R. ; (1999). Carbon dioxide reduction through urban forestry: guidelines for professional and volunteer tree planters. Retrieved from http://www.treesearch.fs.fed.us/pubs/6779
McPherson, E. G., & Simpson, J. R. (2003). Potential energy savings in buildings by an urban tree planting programme in California. Urban Forestry & Urban Greening, 2(2), 73–86. https://doi.org/10.1078/1618-8667-00025
McPherson, G. E., Nowak, D. J., & Rowntree, R. A. ; (1994). Chicago’s urban forest ecosystem: results of the Chicago Urban Forest Climate Project. Retrieved from http://www.treesearch.fs.fed.us/pubs/4285
McPherson, G., Simpson, J. R., Peper, P. J., Maco, S. E., & Xiao, Q. (2005). Municipal Forest Benefits and Costs in Five US Cities. Journal of Forestry, 103(8), 411–416.
Menninger, E., A. (1977). Edible Nuts of the World. Stuart, FL: Southeastern Printing Co.
Miller, H. A., & Lamb, S. H. (1984). Oaks of North America. Happy Camp, CA: Naturegraph Publishers.
Miller, J. H., Chambliss, E. B., & Bargeron, C. (2006, January). Invasive Plants of the Thirteen Southern States. Retrieved March 10, 2008, from http://www.invasive.org/south/seweeds.cfm
Miller, J. H., & Miller, K. V. (2005). Forest Plants Of The Southeast And Their Wildlife Uses (Revised). Athens, GA: University of Georgia Press.
Miller, W. D. (1967). An Annotated Bibliography of Southern Hardwoods. Raleigh: North Carolina Agricultural Experiment Station.
Miller, W. D. (1974). An Annotated Bibliography of Southern Hardwoods, Vol. II. Raleigh, N.C.: North Carolina Agricultural Experiment Station.
Moerman, D. E. (1998). Native American Ethnobotany. Portland, OR: Timber Press, Incorporated.
Moerman, D. E. (2003). Native American Ethnobotany. Retrieved April 23, 2009, from http://herb.umd.umich.edu/
More, D., & White, J. (2005). The Illustrated Encyclopedia of Trees (2nd Ed). Portland, OR: Timber Press, Inc.
Mudge, K., Gabriel, S., & Munsell, J. (2014). Farming the Woods: An Integrated Permaculture Approach to Growing Food and Medicinals in Temperate Forests. White River Junction, VT: Chelsea Green Publishing.
Neel, L., Sutter, P. S., Way, A. G., & Franklin, J. F. (2012). The Art of Managing Longleaf: A Personal History of the Stoddard-Neel Approach. A Wormsloe Foundation Nature Book.
Nelson, G. (1998). The Trees of Florida: A Reference and Field Guide. Sarasota, FL: Pineapple Press.
Nelson, L. S., Shih, R. D., & Balick, M. J. (2007). Handbook of Poisonous and Injurious Plants (2nd ed.). New York: Springer.
Nicholson, S. M., & Clovis, J. F. (1967). Dye Plants and Dye Methods in West Virginia. Castanea, 32(2), 111–116. https://doi.org/10.2307/4032274
Nowak, D. J., & Crane, D. E. (2000). The Urban Forest Effects (UFORE) model: quantifying urban forest structure and functions. Retrieved from http://www.nrs.fs.fed.us/pubs/8420
Nowak, D. J., Crane, D. E., & Stevens, J. C. (2006). Air pollution removal by urban trees and shrubs in the United States. Urban Forestry & Urban Greening, 4(3–4), 115–123. https://doi.org/10.1016/j.ufug.2006.01.007
Nowak, D. J., Crane, D. E., Stevens, J. C., Hoehn, R. E., Walton, J. T., & Bond, J. (2008). A ground-based method of assessing urban forest structure and ecosystem services. Aboriculture & Urban Forestry, 34(6), 347–358.
Nurseries, H. (1992). The Hillier Manual of Trees and Shrubs (6th Ed). Melkham, Wiltshire Great Britain: David and Charles.
Nyland, R. D. (2007). Silviculture: Concepts and Applications (2nd Ed.). Long Grove, IL: Waveland Pr Inc.
Oswalt, S., Thompson, M., & Smith, W. (Eds.). (2009, Rev). US Forest Resource Facts and Historical Trends. US Forest Service. Retrieved from http://www.fia.fs.fed.us/library/brochures/docs/Forest%20Facts%201952-2007%20US%20Metric%20rev072411.pdf
Outland, R. B., III. (2004). Tapping The Pines: The Naval Stores Industry In The American South. Baton Rouge: Louisiana State Univ Press.
Pakenham, T. (2003). Remarkable Trees of the World. New York: WW Norton and Company.
Palm Oil and Tropical Deforestation. (n.d.). Retrieved May 7, 2016, from http://www.ucsusa.org/global_warming/solutions/stop-deforestation/palm-oil-and-forests.html
Pavlik, B. M., Muick, P., & Johnson, S. (1993). Oaks of California. Los Olivos, CA: Cachuma Press.
Peattie, D. C. (2013). A Natural History of North American Trees. San Antonio, Texas: Trinity University Press.
Pellett, F. C. (1977). American Honey Plants (4th ed.). Hamilton, IL: Dadant and Sons.
Pertchik, B., Pertchik, H., & White, W. C. (1951). Flowering Trees of the Caribbean. Rinehart & Company.
Peterson, L. (1978). A Field Guide to Edible Wild Plants of Eastern and Central North America. Boston: Houghton Mifflin.
Petrides, G. A. (1998a). A Field Guide to Eastern Trees: Eastern United States and Canada, Including the Midwest (1st ed., expanded). Boston, MA: Houghton Mifflin.
Petrides, G. A. (1998b). A Field Guide to Western Trees (Second Edition). Boston, MA: Houghton Mifflin Harcourt.
Petrides, G. A. (2005). Trees of the California Sierra Nevada. Mechanicsburg, PA: Stackpole Books.
Phillips, R. (1978). Trees of North America and Europe. (T. Wellsted, Ed.). New York: Random House.
Phillips, R., & Rix, M. (2002). The Botanical Garden: Volume 1 Trees and Shrubs. Buffalo, NY: Firefly Books.
Pojar, J., MacKinnon, A., Alaback, P., Antos, J., Goward, T., Lertzman, K., … Vitt, D. (1994). Plants of the Pacific Northwest Coast: Washington, Oregon, British Columbia & Alaska. Renton, WA ; Vancouver: B.C. Ministry of Forest and Lone Pine Pub.
Porcher, F. P. (1970). Resources of the Southern Fields and Forests. New York: Arno.
Poveda, L. J., Jiménez, Q., & Zamora, N. (2011). Trees of Costa Rica vol. IV. (D. Ávila, Ed.) (First edition (Kindle)). Editorial INBio.
Pratt, H. D. 1944-(Harold D. (1998). A Pocket Guide to Hawai`i’s Trees and Shrubs. Honolulu, Hawaii: Mutual Pub.
Price, -Bargain. (2007). The Garden in Winter: Plant for Beauty and Interest in the Quiet Season (First Edition edition). New york, NY: Rodale Books.
Puettmann, K. J. (2008). A Critique of Silviculture: Managing for Complexity. Washington, DC: Island Press.
Radford, A. E. B., Ahles, H., & Bell, C. R. (1968). Manual of the Vascular Flora of the Carolinas. Chapel Hill, NC: University of North Carolina Press.
Rapp, V. (2002). Life in an Old Growth Forest. Minneapolis, MN: Lerner Pub Group.
Reilly, H. E. (1992). Success with Rhododendrons and Azaleas. Portland, Or: Timber Press, Incorporated.
Rogers, R. D. (2014a). Ancient medicinal remedies: horsetails, ferns, lichens and more.
Rogers, R. D. (2014b). Fairy Wand to Queen’s Cup: The Medicine and Magic of the Order of Lilies. CreateSpace Independent Publishing Platform.
Rogers, R., & Wasser, S. P. (2011). The Fungal Pharmacy: The Complete Guide to Medicinal Mushrooms and Lichens of North America. Berkeley, CA: North Atlantic Books.
Russell, T., & Cutler, C. (2003). Trees: an illustrated identifier and encyclopedia. London: Hermes House.
Russell, T., & Walters, M. (2007). The illustrated encyclopedia of trees of the world. London: Lorenz.
Salley, H., & Greer, H. E. (2005). Rhododendron Hybrids (2nd Ed). Portland, OR: Timber Press, Incorporated.
Sargent, C. S. (1965). Manual of the Trees of North America (exclusive of Mexico) (2d corr. ed, Vols. 1–2). New York: Dover Publications.
Scott, T., L. (2010). Invasive Plant Medicine: The Ecological Benefits and Healing Abilities of Invasives. Rochester, VT: Healing Arts Press.
Seddon, S. A., & Lennox, G. W. (1980). Trees of the Caribbean. London: Macmillan Caribbean.
Shirley, H., L., & Graves, P., F. (1967). Forest Ownership for Pleasure and Profit. Syracuse, NY: Syracuse University Press.
Simeone, V. A., Curtis, B., Barnett, D., & Tankersley, B. (2005). Wonders of the Winter Landscape: Shrubs and Trees to Brighten the Cold-Weather Garden. Batavia, IL: Chicago Review Press.
Simpson, M. G. (2010). Plant Systematics. Burlington, MA: Academic Press.
Sinclair, W., & Lyon, H. H. (2005). Diseases of Trees and Shrubs (2nd Ed). Ithaca: Comstock Publishing Associates.
Smith, D. M., Larson, B. C., Kelty, M. J., & Ashton, P. M. S. (1996). The Practice of Silviculture: Applied Forest Ecology, 9th Edition (9th Ed.). New York: Wiley.
Soltis, D. E. (2005). Phylogeny and evolution of angiosperms. Sunderland, Mass.: Sinauer Associates.
Southgate, D. (1998). Tropical Forest Conservation: An Economic Assessment of the Alternatives in Latin America. New York: Oxford University Press.
Spears, P. (2006). A Tour of the Flowering Plants: Based on the Classification System of the Angiosperm Phylogeny Group. St. Louis, MO: Missouri Botanical Garden Press.
Spichiger, R., Savolainen, R., Figeat, M., & Jeanmonod, D. (2004). Systematic Botany of Flowering Plants: A New Phylogenetic Approach to Angiosperms of the Temperate and Tropical Regions. Enfield, NH: Science Publishers.
Sponsel, L., E., Headland, T., & Bailey, R., C. (Eds.). (1996). Tropial Deforestation: The Human Dimension. New York: Colombia University Press.
Stein, J., Binion, D., & Acciavatti, R. (2012). Field Guide to Native Oak Species of Eastern North America. CreateSpace Independent Publishing Platform.
Sterry, P., & Sterry, P. (2008). Collins complete guide to British trees: a photographic guide to every common species. London: Collins.
Storer, D. P. (1964). Familiar Trees and Cultivated Plants of Jamaica. Macmillan and Co Ltd.
Streich, J. P. (2012). Fat Lighter: Our Southern Longleaf Heritage. Lilburn, GA: CreateSpace Independent Publishing Platform.
Stuart, J. D., & Sawyer, J. O. (2001). Trees and Shrubs of California. Berkeley, CA: University of California Press.
Stupka, A. (1964). Trees, Shrubs, and Woody Vines of Great Smoky Mountains National Park. Knoxville, University of Tennessee Press.
Sudworth, G. B. (1967). Forest Trees of the Pacific Slope. Dover Publications Inc.
Svanberg, I., Sõukand, R., Łuczaj, Ł., Kalle, R., Zyryanova, O., Dénes, A., … Kolosova, V. (2012). {Uses of tree saps in northern and eastern parts of Europe}. Acta Societatis Botanicorum Poloniae, 81(4), 343–357. https://doi.org/10.5586/asbp.2012.036
Swanson, R. E. (1994). A Field Guide to the Trees and Shrubs of the Southern Appalachians. Baltimore: Johns Hopkins University Press.
Symonds, G. W. D. (1958). The tree identification book: a new method for the practical identification and recognition of trees. New York: M. Barrows.
Symonds, G. W. D. (1963). The Shrub Identification Book. New York: M. Barrows.
Tallamy, D. W. (2009). Bringing Nature Home: How You Can Sustain Wildlife with Native Plants, Updated and Expanded (Expanded Edition). Portland, OR: Timber Press.
Tanner, E. V. J. (1977). Four Montane Rain Forests of Jamaica: A Quantitative Characterization of the Floristics, the Soils and the Foliar Mineral Levels, and a Discussion of the Interrelations. Journal of Ecology, 65(3), 883–918. https://doi.org/10.2307/2259385
Tennessee Valley Authority. (1998). Landscaping with Native Shrubs in Utility Rights-of-Way: A Guide to Selecting Native Shrubs for Rights-of-Way Naturalization in the Tennessee Valley. Norris, TN: Environmental Research & Services, Tenn. Valley Authority.
Thomas, P. A. (2014). Trees: Their Natural History (2 edition). Cambridge, U.K. ; New York: Cambridge University Press.
Tiruneh, F. M., & Hager, H. (2011). Edible fruits in the wild and in homegardens: Fruit trees in the wild and in homegardens: species composition, diversity and utilization in Amhara region of Ethiopia. LAP LAMBERT Academic Publishing.
Towe, L. C. (2004). American Azaleas. Portland, OR: Timber Press.
Trelease, W. (1967). Winter botany. New York: Dover Publications.
Trestain, C. G. (1998). Appalachian Willow Work: Re-Establishing a Sense of Place and Developing Concepts of Sustainability (Master’s Thesis). Appalachian State University.
Valentine, J. (2011). Southern Appalachian Celebration: In Praise of Ancient Mountains, Old-Growth Forests, and Wilderness. Chapel Hill: The University of North Carolina Press.
Van Wyk, B.-E. (2005). Food plants of the world: an illustrated guide. Portland, OR: Timber Press.
Van Wyk, B.-E., & Wink, M. (2004). Medicinal Plants of the World: An Illustrated Scientific Guide to Important Medicinal Plants and Their Uses. Portland, OR: Timber Press.
Vanveen, T. (1986). Rhododendrons in America (2 edition). Portland, Oregon: Binford & Mort Pubs.
Wasson, E. (2001). Trees and Shrubs: Illustrated A-Z of Over 8500 Plants. Willooughby, NSW Australia: Global Book Publishing.
Watson, G. (1994). Tree Care Handbook - The Morton Arboretum. Lisle, IL: The Morton Arboretum.
Watson, G. (1998). Landscape Below Ground II: Proceedings of the 2nd Int’L Workshop on Tree Root Development in Urban (1st Ed.). Champaign, IL: Intl Society of Arboriculture.
Watson, G. (2005). Tree Planting. Champaign, lL: Intl Society of Arboriculture.
Watts, M. T., & Watts, T. (1970). Winter tree finder: a manual for identifying deciduous trees in winter. Berkeley, CA: Nature Study Guild.
Way, A. G. (2011). Conserving Southern Longleaf: Herbert Stoddard and the Rise of Ecological Land Management. Athens: University of Georgia Press.
Weakley, A. (2012). Flora of the Carolinas, Virginia, Georgia, northern Florida, and surrounding areas (Working Draft November). Chapel Hill, NC: North Carolina Botanical Garden.
Weakley, A. (2015). Flora of the Southern and Mid-Atlantic States (Working Draft). Chapel Hill, NC: University of North Carolina Chapel Hill Botanical Garden.
Wells, D. (2010). Lives of the trees: an uncommon history. Chapel Hill: Algonquin Books of Chapel Hill.
Whitefield, P. (1998). How to Make a Forest Garden. East Meon, Hampshire, England: Permanent Publications.
Williams, G. (1979). Agrisilviculture for Appalachia: A Bibliography of Relevant Materials Selected from the Library of the International Tree Crops Institute. Gravel Switch, KY: International Tree Crops Institute, U.S.A., Inc., Appalachian Regional Office.
Williams, M. D. (2007). Identifying trees: an all-season guide to Eastern North America. Mechanicsburg, PA: Stackpole Books.
Wink, M., & Van Wyk, B.-E. (2008). Mind-Altering and Poisonous Plants of the World (1st ed). Portland, OR: Timber Press.
Winston, D., & Maimes, S. (2007). Adaptogens: Herbs for Strength, Stamina, and Stress Relief. Rochester, VT: Healing Arts Press.
Wofford, B. E. (1989). Guide to the Vascular Plants of the Blue Ridge. Athens, GA: University of Georgia Press.
Wojtech, M. (2011). Bark: a field guide to trees of the northeast. Hanover, New Hampshire: University Press of New England.
Wolverton, B. C. (1997). How to Grow Fresh Air: 50 Houseplants That Purify Your Home or Office. New York: Penguin Books.
Woodland, D. W. (2009). Contemporary Plant Systematics (4th ed.). Andrews University Press.
Yance, D. R. (2013). Adaptogens in medical herbalism: elite herbs and natural compounds for mastering stress, aging, and chronic disease. Rochester, VT: Healing Arts Press.
Young, J. (1981). Rhododendrons of China. Prepared by American Rhododendron Society and Rhododendron Species Foundation. (L. Chong, Trans.) (First Edition edition). Portland, Or: Binford & Mort Pubs.
Zamora, N. V., Jimenez, Q., & Poveda, L. J. (2004). Árboles de Costa Rica / Trees of Costa Rica, Vol. 3. Editorial INBio.
Zarin, D. J., Alavalapati, J. R. R., Putz, F. E., & Schmink, M. (2004). Working Forests in the Neotropics: Conservation through Sustainable Management? New York: Columbia University Press.
Zomlefer, W. (1994). Flowering Plant Families. Chapel Hill, NC: The University of North Carolina press.
Zucker, I. (1966). Flowering Shrubs. Princeton, NJ: D. Van Nostrand Company Inc.